ENCHYTRAEIDAE 247 



are free from these glands. The chromatophores are at a deeper level, are brown in 

 colour, and branched ; the pigment is in the longitudinal muscular coat and especially in 

 a layer on its inner surface (cf. M. grisea). 



The body-cavity corpuscles are small; many are spindle-shaped, mostly ca. 20 /x in 

 length, though sometimes as much as 30/x; others, about the same length, are much 

 broader and subcircular in form; all are nucleated, the cell body staining lightly with 

 eosin and granular in constitution. 



The postpharyngeal bulbs are the best-developed examples of these enigmatical 

 structures that I have ever met with. There are no salivary glands. 



The septal glands are bulky, and take up a good deal of room in segments iv-vi ; those 

 of vi project back a long way into vii. In the second series of sections to be examined 

 there was in segment vii a separate septal gland, not a part of that belonging to vi. Its 

 connection with the gland in vi was effected by a non-staining stalk or strand (as the 

 septal glands in each successive segment are always connected to those in front), and it 

 passed backwards through a definite opening in septum 7/8 (not merely bulging this 

 septum backwards) so as to occupy also the anterior part of segment viii. Altogether, 

 therefore, the extent of the glands was quite unusual in this specimen. 



The chloragogen cells of the anterior segments contain much brown pigmented 

 matter. 



The dorsal vessel arises in segment xiii, through the whole length of which it extends. 

 From the staining reaction of its contents with eosin the blood presumably contained 

 haemoglobin and was red in life. 



The nephridia possess each a rather bulky anteseptal portion, with a cylindrical 

 funnel and, in addition, an amount of glandular tissue ; the anteseptal (82 or even perhaps 

 90 ju,) is nearly as long as the postseptal (100 /x). The duct is as long as the postseptal; it 

 may be given off from the lower surface of the postseptal some little distance from the 

 hinder end of the organ, when it takes a curved course to the surface; its lumen is 

 dilated, apparently forming a sort of reservoir, where it enters the body-wall. The place 

 of origin of the duct, however, seems to vary ; from the evidence of longitudinal sections it 

 may be given off from the middle of the postseptal, or even from quite near the septum. 



The testes are small organs, each consisting of 4 or 5 cylindrical or slightly club- 

 shaped or pear-shaped lobes surrounded by no obvious capsule ; the type of organ is 

 really that characteristic of the genus Lumbricillus, since the organ is divided down to its 

 base. Segment xi contains a large number of free male cells in various stages of 

 development (morulae, bundles of spermatozoa) — so many that the anterior septum is 

 much bulged forwards, nearly to the level of furrow 9/10; the ends of the testis lobes 

 break up, and the cells become free, much earlier than in most species oi Lumbricillus, so 

 that the testes are here much smaller organs than usually in that genus. 



The male furmel is about zh times as long as broad; it is a stout organ, about half as 

 broad as the available diameter of the segment ; its everted rim or collar is narrow and 

 projects only slightly. The vas deferens, 1 3 /^ in diameter, is a loose coil of no great length ; 

 it reaches the surface through a cleft in the muscular mass which represents the penial 



