ENCHYTRAEIDAE 261 



There are no salivary glands, but a pair of postpharyngeal bulbs arise, as in the 

 genera Marionina and LiimbriciUm, from the recess behind the dorsal pharyngeal plate. 



There is no sudden widening of the oesophagus. In specimens mounted whole the 

 very dark chloragogen investment, beginning in segment v, is remarkable, and is rather 

 characteristic of the species. The chloragogen cells are large and elongated, up to 57 /x 

 in height ; the nucleus is not far from the middle, and the greater part of the interior of 

 the cell is taken up by half a dozen or more large vacuoles, in series or sometimes two 

 abreast (this appears to be the condition which is described for E. biichholzi). The very 

 small (ijLi or less) brown chloragogen particles are numerous in the anterior segments. 



The dorsal vessel begins at the anterior end of segment xiv, just behind septum 13/14 

 (in two sectioned specimens), or (in a third) extends through the whole of segment xiv. 

 The blood stains red with eosin (indicating the presence of haemoglobin), and was 

 therefore red during life; in some of the vessels are red-staining crystals, perhaps 

 crystals of haemoglobin. 



The nephridia present an extremely elongated funnel constituting the preseptal 

 portion, 80 ft in length by 20 /^ in diameter, with long cilia pointing down the tube and 

 others directed outwards from the lip of the funnel. The shape of the funnel appears 

 cylindrical in some cases, more conical, narrowed towards the septum, in others; this 

 depends probably on the plane in which the part is cut; there are two, if not three, 

 nuclei in the funnel, one being in the projecting part of the lip. The postseptal portion 

 is about i50jii long; its apparent width depends on the plane in which it is cut — from 

 65 to 90 /x. The duct, given off from the hinder end of the organ, leads downwards or 

 downwards and forwards. 



The cerebral ganglion (Fig. 12) is as broad as it is long, indented in front and behind, 

 the sides converging forwards ; the breadth at the widest part 

 exceeds that at the anterior end in the ratio 40 : 27. 



The proper testes are quite small, situated anteriorly and 

 ventrally in segment xi, and cut up into a number of irregular, 

 small, ovoid or shortly cylindrical lobes, which in many cases 

 are seen to be surrounded by a delicate membrane. The testes 

 consist only of sexual cells which have not begun to divide to 

 form morulae ; they differ altogether from the lobed (" divided ") 

 testes of the genus Liimbricilhis, where the numerous large club- 

 shaped lobes, each surrounded by a membrane, consist of 

 undifferentiated sexual cells along with all stages — morulae, 



FiET. 12. EfichytYdBus col- 

 spermatids and spermatozoa — of their subsequent development. J, u \ r 



ir tr n r pites; cerebral ganglion. 



In the present species male cells in all stages of development 



are found free in the cavity of segment xi, in such quantities as to bulge septum lo/ii 

 forwards or 11 /12 backwards. What happens therefore is that the enclosing membrane 

 disappears at an early stage, and the lobes of the testis shed their cells into the body- 

 cavity, where their ripening takes place. 



The male funnels are of moderately large size, about 5 or 6 times as long as broad, 



