296 Carl S\ottsberg 



("Panke" is a nonsensical corruption), with about a dozen 

 species along the Andes, 2 in Costa Rica, i in Brazil, and 3 

 in Juan Fernandez, to which the 2 Hawaiian species are most 

 closely related, whereas they have nothing whatever to do with 

 the west Pacific Psetido gunner a. The Paleantarctic character of 

 Gunnera is affirmed by the fairly close relation between Mis- 

 andra and Milligania. 



The distribution of Acaena is equally striking. G. Bitter, the 

 monographer of the genus, recognized 10 sections. Numbers I 

 (13 sp.), IV (i), and VI (i) are confined to the American sec- 

 tor; Number V has 6 species in subantarctic America, i in Masa- 

 fuera, and i in Tasmania; VII has 26 species in South America, 

 I in California, and i in Australia and Tasmania; VIII has 56 

 species in America (the Andes, Patagonia, Fuegia, i extending 

 to South Georgia, Kerguelen, etc.), 2 in the African sector, 3 in 

 New Zealand, and i polymorphic species in New Zealand and 

 the surrounding islands, Tasmania, Australia, and New Guinea. 

 Section II is endemic in the African sector (Cape, i sp.). Thus, 

 this sector contains one endemic section, i endemic species of 

 Section VIII in each of the islands of Tristan da Cunha and New 

 Amsterdam, and endemic subspecies of a Magellanic species in 

 Kerguelen, Crozet, and Prince Edward Island. The Australian- 

 Neo-Zelandic sector has two endemic sections— IX with i species 

 and X with 2 species. In addition, as we have already seen, V is 

 represented by i species (Tasmania), VII by i species, and VIII 

 by 4 species. Evidendy the present center, with respect to both 

 the number of species (97) and of sections (7), lies in extratropi- 

 cal South America. However, 5 sections are found in the Aus- 

 tralian-Neo-Zelandic sector (9 sp.), and 2 in the African sector 

 (4 sp.), and, if we beUeve that the stations in these two sectors 

 are nothing but "branch offices" from the headquarters in South 



