I20 



DISCOVERY REPORTS 



Family TOMOPTERIDAE 

 Genus Tomopteris, Eschscholtz 



The parapodial trunks, i.e. the dorsal and ventral divisions of the feet, are bordered 

 all round by membranous wings or pinnules. 



In Tomopteris, sensu stricto, a tail is almost alvi^ays absent, and also, in the adult, the 

 first pair of chaetigerous appendages. Hyaline glands are usually present; and there 

 are no rosettes. 



There are no hyaline glands 



Hyaline glands are present on the ventral pinnules ... 



Chromophil gland on a level with the tip of the ventral trunk 



Chromophil gland below the ventral trunk 



A large species with the gonad in both rami of the feet 

 A small species with the gonad in the dorsal ramus only 



... T. cavallii 



2 



T. septentrionalis 



3 



T. carpenteri 

 T. planktonis 



Tomopteris (Tomopteris) carpenteri, Quatrefages. 



Augener, 1929, p. 304. 



Occurrence. St. 124 (i); 128 (i); 133 (6); 136 (2); 138 (3); 139 (6); 160 (i); 374 (16); 459 

 (2 juv.); 460 (i); 527 (24); 567 (16); 1148 (3); WS 22 (2); WS 26 (3); WS 35 (5); WS 38 (i); WS 39 

 (I); WS 45 (I); WS 53 (I); WS 53B (i); WS 54 (i); WS 55 (i); WS 536 (5); WS 541 (2); WS 544 

 (3); WS 545 (i); WS 547 (i); WS 548 (i); WS 549 (2). 



Specific characters. A large species measuring up to 70 mm. in length for about 

 35 pairs of feet. The prostomium has no median notch. The neck is short and broad. 

 The cerebral ganglion is transversely elongated and slightly bilobed. There is a single 

 pair of black eyes, often invisible in the adult. The anterior pair of chaetigerous ap- 

 pendages is absent. The second pair is globular at the base and may reach back as far as 

 three-fourths of the length of the body. The feet are conical, and the pinnules oval and 

 rather long. The latter have their origin a short distance before the separation of the 

 foot into its constituent rami. From the third foot backwards there is a conspicuous 

 hyaline gland on the pinnule of the ventral ramus a little above and beyond the apex of 

 the pedal trunk. From the fourth foot there is a very large chromophil gland lying 

 below the apex of the pedal trunk in the ventral ramus. Genital products are found in 

 both rami of the foot. 



Remarks. It is noteworthy that out of all these specimens there is not one that I 

 recognized as a female. I do not claim to have "sexed" every specimen carefully, but 

 in the adult examples the foot usually contains a flocculent white substance which 

 penetrates into both trunks after the division of the foot into its two rami. This sub- 

 stance appears to be sperm, and I have never seen any ova in the feet. In other species, 

 e.g. in septentrionalis, ova are very conspicuous, and I think I should have seen them 

 if they had been present in carpenteri. 



Augener records T. platiktotiis from the Weddell Sea and suggests that previous 

 authors have confused it with carpenteri, to which in the arrangement of the pedal 

 glands it is very similar. It is distinguishable by its very much smaller size — it reaches 



