SYSTEMATIC ACCOUNT l8 5 



Genus Vanadis Claparede, 1870 

 Body elongate. Prostomium with two pairs of small antennae on the anterior border and one single, 

 dorsal, and posterior to these. There are three to five pairs of tentacular cirri. Parapodia with long 

 compound chaetae and pedal lobe with terminal appendage. Proboscis sometimes with two long 

 cirriform appendages and always with small terminal papillae. 



Type species. Vanadis formosa Claparede, 1870. 



Type locality. Gulf of Naples. 



Vanadis formosa Claparede, 1870 



Vanadis formosa Claparede, 1870, pp. 480-4, pi. 10, fig. 3. 



Vanadis pelagica Greeff, 1876, p. 67, pi. 3, figs. 33-4. 



Alciopa longirhyncha Greeff, 1885, pp. 453-5, pi. 14. H- 37- 



Alciopa Krohnii Hering, 1892, p. 738, pi. 4, figs. 1-13. 



Vanadis formosa Apstein, 1900, pp. 8-10, pi. 1, fig. 1-6. 



Vanadis formosa Lo Bianco, 1904, p. 50, pi. 29, fig. 113. 



Vanadis formosa Southern, 191 1, pp. 2-3. 



Vanadis formosa Fauvel, 1916, pp. 65-6. 



Vanadis formosa Mcintosh, 1922, p. 13. 



Vanadis formosa Fauvel, 1923, pp. 205-6, fig. ■j'ja-c. 



Vanadis formosa Mcintosh, 1923, pp. 440-1. 



Vanadis formosa Monro, 1930, p. 81 (in part). 



Vanadis formosa Fauvel, 1932a, pp. 16-17. 



Vanadis formosa Augener, 1934, pp. 217-19. 



Vanadis formosa Monro, 1936, p. 116. 



Vanadis formosa Monro, 1937, p. 268. 



Vanadis formosa Monro, 1939a, p. 348. 



Vanadis formosa Wesenburg-Lund, 1939, pp. 28-32, figs. 20-21, chart 15. 



Vanadis formosa Stop-Bowitz, 1948, pp. 25-6, fig. 17, chart 18. 



Vanadis formosa Stop-Bowitz, 195 1, p. 7. 



Description. This species may measure up to 300 mm. in length by 4-5 mm. wide across the bases 

 of parapodia and 12 mm. across the tips of chaetae, but complete specimens are rare. The two pairs 

 of anterior antennae are digitiform: the single antenna is between the large eyes. The proboscis has 

 two long terminal processes, each carrying a pair of basal ailerons. These ailerons are continuous with 

 dorsal and ventral circlets of papillae; there are six papillae in each of the dorsal and ventral groups 

 and they form a coronet terminal on the proboscis. There are three pairs of tentacular cirri on each of 

 successive segments behind the prostomium, of which the first pair are the longest and are joined to 

 each other by a large basal ceratophore across the ventral surface behind the prostomium. The first 

 two pairs of parapodia are reduced or modified in the mature female, the dorsal cirri becoming 

 voluminous seminal pouches. In the male there is no modification of anterior parapodia but 

 the first two pairs are reduced. In both sexes from the third foot onwards, parapodia are well 

 developed with large foliaceous dorsal and smaller ventral cirri; each foot has a pedal mamelon with 

 a projecting acicula, a cirriform appendage and compound chaetae with short terminal articles. 

 Darkly pigmented segmental glands appear on each foot after the first three, except in the mature 

 females when they appear on the first two modified parapodia. In some specimens which have no 

 seminal pouches on anterior feet, there are well-developed white ventral glands appearing at the 

 twenty-fifth parapodia and continuing up to the end of the body. St0p-Bowitz (1948) has associated 

 this gland with the male. 



