, 74 DISCOVERY REPORTS 



pp. 1 81-182). The third specimen, 9 mm. long for twenty parapodia, is too macerated to allow of 

 specific determination. 



(b) Monro, 1936, p. 121, as T. cavalli from 'Discovery', St. 451, 170-0 m. Monro reported five 

 specimens, nine are present in the B.M.(N.H.), (Reg. no. 1936.2.8. 1000-6). All these specimens are 

 small, between 3 and 6 mm. long for twelve to sixteen pairs of parapodia, and have apical hyaline 

 glands on neuropodial pinnules. Not even in the smallest, 3 mm. long for twelve parapodia, is there 

 a first pair of chaetigers. I consider all these specimens are T. planktonis. 



One of the original localities from which specimens of T. cavalli were first described was in the 

 South Atlantic, off Buenos Aires Province (Rosa, 1907) and I had expected to find it in the present 

 survey. It is, however, very close to T. planktonis, differing only in being without hyaline glands, 

 and in having a mid-ventral chromophil gland. This latter character has been shown (p. 172) to be 

 characteristic of development of T. planktonis and of specimens of T. planktonis which also have 

 fully developed chromophil glands on other feet (Text-fig. 6, p. 173). Moreover all species of 

 Tomopteris normally possessing hyaline glands have a proportion of them pigmented, but occasionally 

 specimens occur with some of these glands on a few parapodia lacking pigment and therefore are 

 difficult to see. In T. planktonis the hyaline glands are often unpigmented and staining of the pinnule is 

 necessary to show up the glands. A negative characteristic of this kind can be very confusing and I 

 doubt whether T. cavalli is a valid species. 



(c) In re-examining Monro's (1936) material of T. septentrionalis from 'Discovery', St. 41 A 

 (B.M.(N.H.), Reg. no. 1936.2.8. 1038) I find it to be T. planktonis. 



General distribution. T. planktonis has been widely reported from the Atlantic at all explored 

 depths: it is circumpolar in Antarctic waters and almost certainly cosmopolitan (see pp. 228-231). 



Tomopteris carpenteri Quatrefages, 1865 



Type locality. The Southern Ocean; 6o° 3' S., o° 6' E. 



Tomopteris carpenteri Quatrefages, 1865, pp. 227-9, P^ 20 > n 8 s - J > 2 - 



Tomopteris carpenteri Ehlers, 1917, pp. 229-31, pi. 12, figs. 9-12 (in part). 



nee Tomopteris carpenteri Mcintosh, 1925, pp. 29-32. 



Tomopteris carpenteri Augener, 1929, pp. 304-7. 



Tomopteris carpenteri Benham, 1929, pp. 191-5, pi. n, figs. 18-23. 



Tomopteris (Tomopteris) carpenteri Monro, 1930, pp. 84-6 (in part). 



Tomopteris carpenteri Hardy and Gunther, 1935, p. 115. 



Tomopteris (Tomopteris) Carpenteri Stop-Bowitz, 1949, pp. 16-17, ^§ s - 6~7* 



Tomopteris (Tomopteris) Carpenteri, Stop-Bowitz, 195 1, p. 9. 



Tomopteris carpenteri, Fauvel, 19536, pp. 4-5. 



Description. This is the largest of all known Tomopteridae, measuring up to 90 mm. long by 

 18 mm. wide for thirty-five pairs of parapodia; normally, however, it is about 55 mm. long by 

 15 mm. wide for thirty-three pairs. In the present survey a larger number of young specimens of 

 this species have been examined and the measurements for a number of these are listed according to 

 size in Table 3. Within the size ranges of these juvenile specimens confusion can arise when separating 

 T. carpenteri from T. planktonis. 



The anterior border of the antennae is entire. In young forms the eyes are always visible on the 

 dorsal surface of the prostomium, but are much less distinct in the adult. A first pair of chaetigers 

 is never present; the second pair may reach back to three-quarters of the length of the body. The 

 hyaline gland is deeply pigmented and first appears in both young and adult specimens on the third 

 neuropodial pinnule, dorsal to the parapodial trunk; it is always conspicuous and is present on each 



