TRICELLARIA 357 



Menipea fuegensis Busk, 18526, p. 21, pis. xix, figs. 1-3; Jullien, 1888, p. 70, pi. vii, figs. 

 8-10, pi. xii, figs. 1, 2; Calvet, 1904, p. 6; Vallentin, 1924, p. 373; Hasenbank, 1932, p. 365. 



Menipea aculeata Busk, 1884, p. 20, pi. iv, fig. 2; Calvet, 1904, p. 6; Marcus, 19216, p. 93; 

 Vallentin, 1924, p. 373. 



Scrupocellaria fuegensis Waters, 1904, p. 24. 



Scrupocellaria bifurcata Kluge, 1914, p. 614, text-fig. 3. 



Menipea patagonica (part) Busk, 18526, pi. xxiii, fig. 1. 



Station distribution. Sub- Antarctic: South Atlantic Ocean, Sts. 51, 53, 652, 1230, 1902, 

 WS72, WS79, WS84, WS85, WS87, WS88, WS95, WS220, WS222, WS244, WS245, 

 WS 838, WS 847, WS 871. 



Geographical distribution. Chile (Busk); Magellanic Region (Busk; Jullien; Waters; Calvet; 

 Hamburg Museum, B. 1205; Discovery); Patagonian Shelf (d'Orbigny; Busk; Calvet; Vallentin; 

 Discovery) ; off Patagonian Shelf down to 339 m. (Discovery) ; Kerguelen (Busk ; Kluge ; Hasenbank ; 

 Monod and Dollfus); South Georgia (Calvet); Bouvet Island (Hasenbank); Campbell Island 

 (Marcus) ; Auckland Island (National Antarctic Expedition) ; New Zealand (Hamilton ; 97 . 5 . 1 . 265 ; 

 99-7- 1-734)- 



The synonymy of this species has been discussed by Waters, Marcus and Harmer. 

 Waters considered that he had no proof that there were not three distinct species. 

 Marcus accepted Tricellaria aculeata (Busk) as the form intended by d'Orbigny and 

 thought that T. fuegensis (Busk) was probably the same thing, but was not satisfied 

 about T. fuegensis (Jullien). Harmer regarded them all as representing a single variable 

 species, and after examining the Discovery collection and the specimens in the Busk 

 and other collections in the British Museum I agree with him. 



In the one small colony from Challenger St. 314 zooecia can be seen in which the 

 scutum is simple and unbranched, bears two, three or four points, or is completely 

 absent ; the lateral avicularium may be present or absent and varies in size ; the frontal 

 avicularium is occasionally absent and the distal spines may be stout or slender, long or 

 short, varying in number from two to four. The zooecia are more delicate and slender 

 than in the specimen described by Busk in 1852, but other specimens are more or less 

 intermediate. I therefore regard T. aculeata (Busk) and T. fuegensis (Busk) as synony- 

 mous. Marcus was doubtful whether there might not be a difference in the cryptocyst 

 of these two forms, but I have not detected any. 



Apart from features that have already been shown to be variable, such as the scutum 

 and spines and the presence or absence of frontal avicularia, the chief feature mentioned 

 by Waters as distinguishing T. fuegensis (Jullien) from T. fuegensis (Busk) is the large 

 number of zooecia in the fertile internodes of the former (3-14 in the figure). Ovicells 

 are completely absent from Busk's type-specimen but were on short internodes in 

 Waters 's own specimen which agreed with T. fuegensis Busk in other respects. In the 

 present material the number of the zooecia in the fertile internodes varies from three 

 to fifteen. Those colonies with the longest internodes (namely, those from St. 53, in 

 which the full range from three to fifteen is found, and some from the Falkland Islands, 

 24 . 9 . 1 . 1 , with a maximum internode of ten zooecia) have certain features in common 

 with Jullien 's figure. The non-fertile internodes consist of relatively long, stout zooecia, 



