52 T. F. GOREAU AND AV. D. IIARTMAN 



duced in reef communities. This fraction may also have important 

 lubricating and fluidizing properties which speed the drainage of 

 skeletal sediments from the reef. 



Acknowledgments. This work was supported by the National Science 

 Foundation and the United States Navy Office of Naval Research. 



The authors record their gratitude to E. A. Graham, D. L. Fraser, T. A. 

 Smith, D. A. Catt, R. L. Walker, and L. M. Passano, who were members 

 of the scientific diving team at various times and thus made the under- 

 water studies possible. We also thank Professor J. W. Wells for supplying 

 the identification of the coral Agoricia undata, and Dr. K. E. Chave for 

 critical discussion. The underwater photographs are by T. F. Goreau. 



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