THE GENETICS OF HABROBRACON JUGLANDIS ASHMEAD 



with colorless ocelli. V/hite is' of normal 

 viability and fertility. In combination with 

 shot-veins, to be described later, white con- 

 sistently shows small red flecks scattered in 

 the posterior and ventral part of the eyes. 

 White shot-veins is, therefore, known as varie- 

 gated (Fig. 3). 



CARROT, wh^ (eyes). In March, 1932, mutant 

 males with carrot eyes appeared. This eye color 

 closely resembles orange. White is almost dom- 

 inant to carrot. The white-carrot heterozy- 

 gote has a yellowish cast by which it can be 

 accurately separated from white. Both of these 

 alleles are of normal viability and fertility. 



SPECKLED, Sk (eyes). This factor is due to a 

 dominant gene and results in numerous small, 

 bright red flecks of pigment in a white eye. 

 Preliminary work indicates that the specks are 

 not on the cornea but lie in the soft tissue 

 beneath. They do not appear in the pupal eye 

 until about the third day of pupal life. Speck- 

 led varies in its expression in the two sexes) 

 and is sensitive to temperature. At 30° C, a 

 few specks are visible in the eyes of the fe- 

 males, which vary among themselves in the de- 

 gree of speckling. At room temperature, specksi 

 are usually lacking in female eyes. In males- 

 at 30° C, the degree of speckling is very pro- 

 nounced both as to depth of coloration and num- 

 ber of specks. Although specks are scattered 

 at random throughout the compound eye, there is 

 a denser speckling in the ventral half of the 

 eye. Males, too, vary among themselves in the 

 intensity and the number of specks. At room 



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