118 CHARLES R. STOCKARD 



rotation of the wrist joint. The dorsal surface of the foot 

 loses its forward position and is rotated outward laterally. 

 The entire foot abducts widely, with the wrist, as a conse- 

 quence, knocked in medially and almost contacting' the simi- 

 larly bent wrist of the other front leg. 



Leg skeletons <>]' the F. 2 bassethound-bulldog. Many com- 

 plicating factors are involved in the structural patterns of 

 the bassethound-bulldog hybrids and these will be discussed 

 in various connections in the further chapters of this study. 

 As we have seen, the achondroplasic short legged condition 

 is limited to an entirely separate or independent growth 

 reaction and follows the same genetic basis in these hybrids 

 as was found for the two foregoing breed crosses. In the F 2 

 generation of bassethound-bulldog hybrids, we find individual 

 sports toward dwarf size and others in an opposite direction 

 toward gigantism. These conditions will be discussed later, 

 but some attention to leg size is necessary at this time. Many 

 of these complicating growth reactions more or less obscure 

 the breed type of bone inheritance which stands out so clearly 

 in the previous crosses. Even so, there are definite indica- 

 tions of inheritance of bull typed and hound typed bone. 



Plate "21 shows the right front leg skeletons from six F 2 

 bassethound-bulldog hybrids and, for comparative purposes, 

 the leg of an F x (at the light end). The first three skeletons 

 are extracted recessive long legs with no achondroplasic 

 symptoms, yet it is quite evident that they differ greatly in 

 size and type. The first skeleton is from a large male F 2 

 animal which was definitely oversized when compared with 

 either of the parent stocks and weighed almost 50% more 

 than his litter mates; he was rather like a St. Bernard dog 

 in living appearance as well as in skeletal type. Comparing 

 the leg skeleton from this animal, 1312 c?, with the third and 

 fourth skeletons which came from two litter brothers, it is 

 seen to be much larger, with heavier bone, than either of 

 the other two. This rather St. Bernard typed bone arises in 

 the F 2 generation as a neomorph, probably due to a strange 

 and new genetic complex largely different in type from that 



