104 



Fishery Bulletin 90(1). 1992 



(1952). Eighty percent of the females that we classified 

 using gross anatomical criteria were also classified as 

 active or inactive using histological criteria. The results 



were compared to determine the accuracy of identify- 

 ing active and inactive females by gross anatomical 

 classification. 



ALL FEMALES 



Before spawning; N - 949 

 During spawning: N - 1.658 



HISTOLOGICAL 

 CRITERIA 



ACTIVE FEMALES 



Capable of spawning now or in near future. 



ACTIVE - MATURE 



Advanced yotked oocytes preseni. 

 Before 61 4% 

 During: 39.4% 



NONSPAWNING 



No hydrated oocytes. 



No postovulatory foilicle&. 



Before: 60.4% 



During: 33.3% 



SPAWNI^4G 



Hydraled oocytes, or post- 

 ovulatory follicles are present 



Before: 1.0% 

 During: 6.1% 



T 



NO ATRESIA 



No a atresia of advarx»d 

 yolked oocytes. 

 Before: 38.4% 

 During: 16.4% 



MINOR ATRESIA 



a atresia of advarx:ed yolked 



oocytes Is between 1% arxi 49%. 



Before: 22.0% 



During: 16.9% 



NO ATRESIA 



No a atresia of advanced 



yolked oocytes. 



Before: 0.4% 



During: 4,0% 



MINOR ATRESIA 



a atresia of advanced yolked 

 oocytes Is between 1% arxJ 49% 

 Before: 0.6% 

 During: 2,1% 



INACTIVE FEMALES 



Not capable of spawning now or In near future. 

 Before 38.6% 

 During: 60.6% 



IMMATURE 



Unyolked oocytes 



No atresia. 



Before: 4.7% 



During: 12.2% 



UNCERTAIN MATURITY 



No advarx»d yolked oocytes. 

 Before: 31.3% 

 During: 43,1% 



I 



INACTIVE - MATURE 



Before: 2.5% 

 During: 5.3% 



ATRETIC UNYOLKED 



Unyoll^ed oocytes preseni. 

 With atresia. 

 Before: 21 .0% 

 During: 31 .0% 



I 



EARLY YOLKED 



Early yolked oocytes present. 

 May ftave atresia. 



Before: 10.2% 



During: 12.1% 



MAJOR ATRESIA 



Advanced yolked oocytes 

 present, a atresia of 



advanced yolked Is > 50%. 

 Before: 2.5% 

 During: 2.0% 



POSTSPAWNING 



Postovulalory 



lollldes preseni. 



Before: 0% 



During: 3 3% 



WITH ALPHA 



Only a atresia of 



unyolked preseni. 



Before: 15.5% 



During: 17.3% 



WfTH BETA 



^ atresia must be preseni; 



a of unyolked may be preseni. 



Before: 5.6% 



During: 13.8% 



WITH ALPHA OR NONE 



Only a atresia of early yolked 

 present. Or no atresia present. 



Before: 4.7% 



During: 1.6% 



WTTH BETA 



^ atresia must be present; 



a of early yolked may tM present. 



Before: 5.4% 



During: 10.1% 



WriH ADVANCED 



Advanced yolked oocytes 

 present a atresia of 



advanced yolked is > 50% 

 Before: 0.0% 

 During: 0.8% 



WITHOUT ADVANCED 



No advanced yolked 

 oocytes present 



Before: 0.0% 

 During: 2.5% 



Figure 1 



Dendrogram illustrating hierarchial classes of histological criteria of active and inactive ovaries of Dover sole Microstomiis paciflcus. 

 Percentages of females (California and Oregon combined) in each class and subclass taken before (November-December) and during 

 (January- May) the spawning season. 



