Feeney: Post-yolksac development of two southern California sculpins 467 



10(9) Nape pigment present (several melanophores); dorsal gut pigment not in 



bands Oligocottus/Clinocottus type D 



(= C. recalvus (Morris 1951) or C. globiceps or 0. maculosus (Washington 1986). These 

 specimens are most similar to larger C. recalvus from same locality; however, they share 

 characters with all three species (e.g., otic capsule pigment) or with one species (e.g., nape 

 bubble like 0. maculosus). 



Nape pigment absent (or 1 melanophore occipitally); dorsal gut pigment in 



bands Leptocottus armatus 



11(9) Nape pigment present 12 



Nape pigment absent (rarely present in Orthonopias triacis and flexon Chitonotus pugetensis) ... 19 



12(11) Anterior gut pigment present; ventral gut pigment present 13 



Anterior gut pigment absent; ventral gut pigment absent 14 



13(12) Myomeres 40-42; postanal ventral melanophores ~37-38; pigment on 



snout Paricelinus hopliticus* 



Myomeres 27-30; postanal ventral melanophores <15; pigment on snout 



absent Enophrys taurinal 



(= undescribed; only one specimen collected, similar to E. bison description (Richardson and 

 Washington 1980) except for scattered ventral gut pigment) 



14(12) Postanal ventral melanophores 5-14 15 



Postanal ventral melanophores 15-33 16 



15(14) Preanal myomeres 7-9; head pigment usually absent; gut pigment light and 



scattered Oligocottus synderi 



Preanal myomeres 10-12; head pigment present; gut pigment 



heavy Oligocottus/Clinocottus type B 



(= undescribed; looks much like C. analis but has only 10-14 PAVM) 



16(14) Nape melanophores <6 17 



Nape melanophores >6 18 



17(16) Postanal ventral melanophores 15-21; dorsal gut pigment light to 



moderate Clinocottus embryum * 



Postanal ventral melanophores 21-33; dorsal gut pigment heavy Artedius harringtoni 



18(16) Head pigment absent; dorsal gut pigment moderate; nape melanophores 



~18 Oligocottus/Clinocottus type C 



(= undescribed; no head pigment, 17-31 PAVM; lighter pigment than C. analis, may be 

 0. maculosus) 



Head pigment present (except in smallest larvae (<3.0mm)); dorsal gut pigment heavy; 



nape melanophores < 17 Clinocottus analis 



19(11) Postanal ventral melanophores 7-18 20 



Postanal ventral melanophores ^24 21 



20(19) Ventral gut pigment absent; preanal myomeres 8-10 Artedius creaseri 



Ventral gut pigment present; preanal myomeres 11-12 Cottus asper 



