248 LEAF-ANATOMY OF SdiVOLA CRASSIFOF.t A. 



as 1876, Vesque{5) published an account of the anatomy of 

 Goodenia ovata, which, however, was restricted to an investiga- 

 tion of the anomalous stem-structure. With this, he compared 

 the stems of Scievola crassi folio,, S. Pfuniieri, S. microca7'pa, and 

 S. spinescens, but there is no investigation of leaf-anatomy in 

 any of these species. 



In the Systematic Anatomy of the Dicotyledons (i., p. 471 ), 

 Solereder gives a Ijrief account of the leaf-structure of the 

 Goodeniace*, based upon his own observations of Goodenia 

 ovata, Selliera radicans^ Screvola Plumieri, and Damjnera 

 Brownii, and upon those of Briquet in the case of Bninoiiia. 

 He records that the stomata may occui* upon both surfaces, or 

 upon the lower surface only, and that the number of subsidiary 

 cells, when these are present, is four, two of which are placed 

 parallel to the pore, the other two being applied to the narrow 

 ends of the guard -cells. In form, the leaf is found to be typically 

 bifacial, as exemplified by Daniprera Brotvnii^ but is sometimes 

 centric, as in SccEvola Vluinieri. 8ilicified groups of cells are 

 common, in the formation of which certain epidermal and the 

 adjacent mesophyll-cells take part. As well as the ordinary 

 clothing-hairs, which are characteristic of most members of the 

 Order, peltate, glandular hairs have been (observed in Goodenia 

 ovata and Sccevola Fhtmierl. These are described as being 

 " shortly stalked, peltate glands, the shield of which consists of 

 a few cells separated by radial walls." 



The above account is a summary of our present knowledge of 

 leaf-anatomy in the Goodeniacea'. It is interesting to note that, 

 in none of the plants hitherto investigated, has any copious, 

 epidermal secretion been described. In *S'. crassij'olia, however, 

 the buds and young leaves are coated on both surfaces with a 

 thick layer of sticky resin, which is secreted by numerous, 

 stalked, glandular hairs. 



The secretion of mucilage or resin, or both together, has long 

 been recognised as a method of bud-protection. Groom (2) has 

 given an interesting account of the mucilage- and resin-secreting 

 organs of buds. 'iliis work is restricted to those plants whicli 

 bear "colleters" or "villi" on the stipules. These " colleters,"' 



