BY R. J. TILLYARD. 



311 



The following Table will exhibit, in a concise manner, the 

 differences in the c(jndition of the wing-coupling apparatus studied 

 above : — 



Form of the Colpling-appakatus in the Archetype of the 



Lepidoptera. 



Reviewing all the above evidence, it becomes clear that the 



oldest existing form of coupling-apparatus, viz., that of the 



Micro2)terygidce, is already specialised in a direction from which 



* It is not suggested here that the evolution of the amplexiform type in 

 the iSaturniina and Lasiocanipina has been along the same lines as that of 

 the Papilionina. In the Lasiocanipina, at any rate, a studj'^ of the con- 

 ditions to be seen in the Drtpanido' would suggest a direct descent from 

 Eupterotid-like ancestors with the frenate wing-coupling apparatus fully 

 developed. Thus the Papilionina are probably of remoter origin than 

 either the Saturniina or the higher families of the Lasiocanipina. 



