Pammel — A7iat07nical Characters of Seeds of Leguminosae. 219 



endosperm mucilaginous. Cotyledons with one or more rows 

 of palisade cells ; chlorophyll grains in the cotyledons. 



Coronilla. 



EuHEDTSAREAE. Cuticle irregular, cuticularized layer 

 sharply defined; endosperm reduced. Starch present. 



IIedysaru7n. 



Aeschynomeneae. Cuticularized layer narrow; osteo- 

 sclerids I-shaped ; nutrient layer much compressed; endo- 

 sperm present, more or less evident Aeschynomene. 



Adesmieae. Malpighian cells absent, short, or well devel- 

 oped ; osteosclerids well developed or scarcely differentiated 

 from the nutrient layer; endosperm reduced. Palisade 

 parenchyma of the cotyledons wanting or well developed. 

 Reserve material consists of fat, starch, and aleurone grains. 



1. Malpighian cells wanting Chapmannia. 



1. Malpighian cells present (2). 



2. Malpighian cells short and thin-walled (3). 



2. Malpighian cells long and thick- walled Zornia. 



3. Cotyledons contain starch ,Arachis. 



3. Cotyledons do not contain starch Stylosanthes. 



Desmodieae. Cuticularized layer inconspicuous, light line 



close under the cuticle; osteosclerids I-shaped; endosperm 

 present, the amount variable. Cotyledons with several rows 

 of palisade cells; starch absent. 



1. Endosperm usually copious Lespedeza. 



2. Endosperm less copious Desmodium. 



Vicieae. 



Malpighian cells well developed, surface usually irregular, 

 cells contain one or more chromatophores; osteosclerids 

 I-shaped, large with longitudinal pore-canals; nucellus more 

 or less evident; endosperm reduced. Epidermal cells of the 

 cotyledons much smaller than those of the parenchyma, pal- 

 isade cells absent. Reserve food material consists of fat, 

 starch, and aleurone grains. It is diflScult to differentiate 

 Vicia and Lathyrus. 

 1. Cuticle slightly irregular on the surface Pisum, 



1. Cuticle very irregular on the surface (2). 



2. Lateral walls of Malpighian cells undulated Cicer. 



2. Lateral walls not undulated Vicia and Lathyrus. 



Phaseoleae. 



Cuticularized layer inconspicuous, Malpighian cells usually 

 long; osteosclerids usually I-shaped; mycotic layer evident, 

 or reduced; endosperm nearly wanting. Epidermal cells 

 much smaller than the underlying cells, palisade cells usually 



