138 Acarology 



nymphs have a leathery, wrinkled, granulated integument, which is 

 either mammilated or has tubercles. The capitulum in the adults and 

 nymphs is either subterminal or distant from the anterior margin; in 

 larvae it is subterminal or terminal. In depleted adults and nymphs 

 especially, the capitulum is in a more or less marked depression (cam- 

 erostome). Articulations of the palpi of all stages are free (never 



Figure 103 Antricola coprophiliis (Mcintosh), 1935. Ventral view of female 

 (left) and Argas persicus (Oken), 1818. Dorsal view of female (right). (After 

 Cooley and Kohls 1944) 



fused). Porose areas are absent in both sexes. When present the eyes 

 are placed on the supracoxal folds. Spiracles in adults and nymphs 

 are usually placed anterior to coxae iv. Pulvilli are usually rudimen- 

 tary or absent in adults and nymphs but sometimes well developed 

 (functional) in larvae. Nymphal stages are plural and the number 

 variable. 



Genera: 



1. Argas Latreille, 1795 



Type. A earns refiexiis Fabricius, 1794 



2. Antricola Cooley and Kohls, 1942 



Type. Ornithodoros coprophilus Mcintosh, 1935 



3. Ornithodoros Koch, 1844 



Type. Argas savignyi Audouin, 1826 



4. Otobius Banks, 1912 



Type. Argas megnini Duges, 1884 



Discussion: The Argasidae are world-wide in distribution and occur 

 on a wide variety of hosts. Snakes, turtles, many birds, and at least 



