LARVAL forms; OF C'OLEOPTFRA 



14. Laciiiia present. (Ocelli absent; urogoniplii jointed, Avitli basal 



joint very long, the five distal joints small.) 



Sphodrinae {Spliodnis)-^ 

 Lacinia absent 15 



15. ]Meso- and metathorax convex laterally; urogomphi much 



longer than tenth segment, terete, and nodose. (Galea "with 

 the proximal joint twice as large as the distal ; six ocelli.) 



Broscmae ( Brosc us ) 

 Meso- and metathorax subparallel laterally ; urogomphi as long 

 as or shorter than tenth segment, rather flat and membra- 

 nous beneath, either very short, almost triangular, and dis- 

 tant at base, in Dyschirius, or fairly long, with parallel 

 sides, and united at base, in Cllvina. (Six ocelli or none.) 



Dysrhiriinae (Dyscltirius and 

 Clivina) 



16. Legs fossorial ; body with subparallel sides ; prothoracic spiracle 



ver}^ large, first abdominal spiracle large, the rest of normal 

 size. (Sternal sclerites of abdomen closely adjacent; either 

 six ocelli or none ; urogomphi either smooth and curved 

 toward each other, in Pasiinarhus, or nodose and straight, in 



ISc(iyifes) Scaritinae {Scarites and Pasi- 



machiis) 

 Legs ambulatory or rasorial ; body fusiform ; prothoracic spira- 

 cles of normal size. (Six ocelli) 17 



17. Urogomphi either with short, setose ramuli, in Elaphrus, or 



thick, cylindrical, and beset with numerous setae, in Ble- 

 thisa; four anal prominences. (Nasale extended into a short 

 horn; with or without lacinia.) 



Elaplirinae {Elophrus and Ble- 

 fhiso) 

 Urogomphi terete, nodose or with several setae; two or no 

 anal prominences 18 



18. Penicillus absent ; lacinia absent ; claws slender and not strong ; 



setae of femora fine Pairohinae {Pairohus)^^ 



Penicillus present ; lacinia present or at least represented by a 

 strong seta ; claws of normal strength ; setae of femora mod- 

 erately strong. (Anterior margin of nasale varying much 

 according to genus; six ocelli, except in the blind Sphodrop- 



2® This subfamily differs considerably from the Pterostichinae in 

 which, according to the characters of the imagines, its only genus, 

 Sphodrus, has been placed between Pterosfichits and Laemostenus, 

 but it appi-oaches, together with the Broscinae, the great and well- 

 defined bembidiine association. 



^"^ See explanatory remarks on page 19. 



22 



