LARVAL FORMS OF C'OLEOPTERA 



Body depressed ; liead porrect, laterally parallel ; mala subtri- 



aiigiilar Sfaphi/linidae - Hahrocerinae (0 / is - 



lli(i()-us and HahrocerusY^ 



Stenine association : 

 22. Urogomplii long and two-jointed; antenna more than twice as 

 long as head; lignla bilobed; six ocelli on each side. 



Staph ylinidae-Steninae (pi. 17 A. C, 



Urogom))hi absent or small and immovable ; antenna not longer 

 than head ; ligiila absent ; less than six ocelli, sometimes no 

 ocelli 25 



Staplnjlinine association : 

 2."). Legs strong, fossorial; urogomphi one-jointed, thick, sansage- 



like Staph i/linidae-Th inopininae (pis. 15 



B and 18 A, D, E, CI. H-J) 

 Legs cnrsorial ; nrogomphi two-or three-jointed 24 



24. Ocelli, four or less on each side. 



Staph ylinidae-Staph i/Iin inae (Xa n th o- 

 linus with one ocellus, Othius with 

 none, Quedius in many species with 

 club-shaped or capitulate setae, 

 Staph ylinus, Phdonthus, and other 

 genera. ) 



Ocelli, five or six Staphiilinidae-Pacderinae (pi. 18 B, 



C, F) 



Pselaphid association: 



25. Terga ex])anded laterally, body oval 26 



Terga not expanded. (Antenna not club-shaped.) 



Pselaphidae*° {Batrisodes and Euplec- 

 tus) (pi. 19 E-J) 

 2(). Tergal shields smooth, with simple hairs; antenna Avith second 

 joint very large and clnb-shaped. 



Sc\jdmaenidac*^ {Scydmaenus and Eu- 

 micriis) (pis. 16 A-E and 19 A-D) 

 ^^ The larvae of Olisthaerus, of which two European species are 

 completely described and figured by Saalas (Uunio Saalas p. 69) 

 agree in every character with Hahroccrus (schwarzi Horn) and the 

 two genera constitute together a subfamily that comes close to the 

 staphylinine association having a porrect head and antennae in- 

 serted dorsal ly near the anterior margin of the head. The sub- 

 family also approaches the Piestinae and Oxytelinae, thus forming 

 a remarkable link between the more primitive and the highly trans- 

 formed and advanced Staphylinidae. 



;50 



