Subfamily Glavigerinae 



The Glavigerinae usually have the following combination of characters. 

 The head is longer, much longer than wide, and subcylindrical; very rarely 

 is the head subquadrate, or as wide as long. A median gular fovea is present; 

 the pair of vertexal foveae is present, often small, but at times absent 

 (Adranes). The eyes may be well-developed to rudimentary in the same genus 

 (Fustiger) , or absent {Adranes, Claviger) . The mouth-parts are always rudi- 

 mentary — in fact, this is one of the best characters in defining the subfamily. 

 The mandibles are usually slender, terminating in a blunt point, and lacking 

 teeth. The maxillae are vestigial, provided with a distinct brush of setae, used 

 m scraping their specialized food. The maxillary palpi appear to be of a single 

 segment, or at most two segments, with the distal piece usually pedunculate, 

 and terminating in setae rather than a palpal cone. This palpomere is elongate- 

 truncate (Claviger) or subspherical to ovoidal [Claviger odes) . Mentum very 

 large. Labrum very large. Thus these small mouth-parts (mandibles, maxillae, 

 labium) are only studied with precision from slide-mounts of dissected parts. 

 In life these oral organs are relatively hidden by the mentum and labrum, the 

 oral aperture being often a transversely oval, sharply delimited area with the 

 mouth-parts somewhat recessed. This means that apprehension of living and 

 struggling prey by toothed mandibles is impossible, rather the food must 

 be passive, and nourishment obtained either by piercing with the mandibles, 

 then sucking and licking the exuded liquids, or scraping, brushing, and licking 

 a surface. 



The antennae have but one feature in common, namely the distal anten- 

 nomere is, as far as I know, more or less sharply truncate, with the circular to 

 oval truncation provided with erect setae. This is not usual in the Pselaphinae. 

 The number of antennomeres is highly variable. Thus the antennae have two 

 segments {Mastiger, Articerus, Disarthricerus) , three (Adranes, Fustiger, 

 Apoderiger), four (Rhynchoclaviger, Radama, Diartiger), five (Claviger odes, 

 Paussiger, Microclaviger) or six (Claviger, Claviger opsis, Pseudacerus) to 

 mention but three genera in each type. This variation in antennomeres is an 

 excellent character for separation of groups of genera within the subfamily, 

 but is not valuable as a phylogenetic feature for separating Glavigerinae from 

 Pselaphinae, since this variation is surpassed in Goniacerini. Thus clavigerines 

 have a variation of from two to six segments, while Goniacerini has a variation 

 of from five to eleven segments. The first clavigerine antennomere is invisible 

 from above, deeply inserted into the acetabulum at each side of the front. 



Elytra with the apical margins bearing a trichome of long, tortuous, golden 

 setae near each external angle; only rarely are these trichomes absent — they 

 are badges of the symphile. 



(350) 



