66 Frogs of the Okefinokee Swamp 



the center of Scaphiopus (S. hammondi, S. hurteri, S. couchii) is southwestern 

 and that *S. holbrookii came from *S. couchii? Or are the two diverse Pseudacris. 

 lines: P. nigrita (clarkii) and P. ornata the center of all the others unless 

 it be P. ocularis? Or is Gastrophryne carolinensis the one eastern fringe 

 species from a Mexican center with several species? 



5. Or must we think of some of the southwestern forms as outliers from 

 an eastern center? Hyla arenicolor a sole representative of Hyla femoralis 

 and Hyla versicolor? Or is Hyla regilla the one western approach to Pseu- 

 dacris species? 



6. Are Hyla gratiosa, Pseudacris ocularis, Bufo quercicus, Rana heck- 

 scheri, archaic or primitive relicts of an old faunal center in central Florida 

 or elsewhere in the southeast comparable to the peninsular forms such as 

 Neoseps, Rhineura and several snakes? 



7. Or have some of these had ingress quite recently or accidentally, 

 possibly like E. ricordii, geckos, etc.? 



8. Are no certain Nearctic or Holarctic types like Rana sylvatica, or 

 some Pacific Ranas or Ascaphus truei represented in the Okefinokee region? 

 Just as in mammals, birds, etc. slight if any true upper Austral or Transition 

 Zone forms appear. 



9. Are the Atlantic coast forms like Bufo fowleri, Hyla andersonii, Rana 

 virgatipes absent or scarce just as Chrysemys picta, Clemmys muhlenhergii, 

 etc. are absent, Clemmys guttata very rare, etc., i.e. is the Atlantic coastal 

 influence attenuated almost to nothing and Sabalian conditions predominant? 



10. These queries might be expanded with no positive statements made 

 but enough herein appear to show the bases for these imaginative indul- 

 gences or speculations. 



SUMMARY OF AFFINITIES 

 In a sentence we might venture the following tentative assertions regarding 

 each species of this study. 



Scaphiopus holbrookii falls in relationship with Scaphiopus couchii, S. 

 hurteri and less so with S. hammondii. 



Bufo quercicus belongs near some western or southwestern forms such 

 as Bufo compactilis and others. 



Bufo terrestris may be of the Bufo fowleri, B. americanus group or 

 related to Biifo valliceps. 



Acris gryllus, the most aberrant Hylid, possibly more Ranid (see 

 hind femur like R. grylio or R. virgatipes or tadpole) but possibly near 

 Hyla crucifer or Pseudacris ocularis. 



Pseudacris nigrita, much Uke P. triseriata, P. feriarum, etc., and pos- 

 sibly of a different line from Pseudacris ornata. 



Pseudacris ocularis may be as related to Hyla crucifer or Acris gryllus 

 as any of the Pseudacris nigrita assemblage. 



Hyla andersonii probably falls close to Hyla cinerea group or possibly 

 near Hyla squirella. 



