396 Frogs of the Okefinokee Sivamp 



In 1928 we collected many intermediate sizes between transformation 

 and 82 mm., but they were mislaid. These with the last 5 live intermediates 

 of 1922, might have outlived the possible growth groups. With our meager 

 material we might compare it with the possibly comparable Raria grylio. 

 We might postulate: 30-49 mm., average 35.5 mm., transformation; 56 mm., 

 I year old; 82 mm., 2 year old; 91, 95, 95, 97 mm. cf s, 102 mm. 9 , 3 year 

 olds; 113 mm. cf, 4 year old; 125 mm. 9 , 5 year old. 



ENEMIES 



On June 8, 1928 we found a large southern water snake (Natrix sipedon 

 fasciata) on the mud flats where transformed frogs were or at times it was 

 coursing in water where countless tadpoles were. The pied water snake is 

 also an enemy of these tadpoles and young frogs. On June 9, 1928, in the 

 clay hole where the youngest tadpoles were numerous large mouthed black 

 bass lived. These doubtless feed on the tadpoles. At the same time Florida 

 grackles were apparently feeding on the transformed frogs along the edges of 

 the pond. 



At the type locahty we took one transformed frog with all of left leg gone 

 below femur. Another had the left leg completely severed and the toes clean 

 cut from the right foot. In this hole we took Sternotherus minor. The above 

 work is quite clearly the work of turtles or possibly fish. 



AFFINITIES 



The original diagnosis is pertinent in this instance. 



Diagnosis. Like Rana grylio and Rana catesbeiana, it has no dorso-lateral 

 fold and no phalanx of the fourth toe is totally free of web; third toe in 56 mm. 

 specimen is i to 3 mm. shorter in Rana heckscheri than in the other two species 

 of bullfrog or 3 to 6 mm. shorter in 82 mm. specimen or 6 to 9 mm. shorter in 

 95 mm. specimen; third toe 3.8 (95 mm.)-3.56 (82 mm.)-3.3 (56 mm.) in 

 length (snout to vent) in R. heckscheri while 2.7 to 3.1 (95-56 mm.) in R. 

 grylio and R. catesbeiana; third toe 1.6-1.7 in fourth toe in R. heckscheri 

 while 1. 2 -1. 5 in fourth toe in the other two species; first finger decidedly 

 longer than second, while in the other two species it is usually shorter or 

 sometimes equal; first, second, third and fifth toes shorter than corresponding 

 toes of R. grylio and R. catesbeiana; fourth finger 8.6 (95 mm.)-8.2 (82 mm.)- 

 8.0 (56 mm.) in length (snout to vent) while 6.0 to 6.3 (95 mm.)-6.8 to 7.4 

 (82 mm.)-5.6 to 7.1 (56 mm.) in the other two specimens; internasal space 

 less than upper eyelid width, i. 07-1. 43 in it while .85-1.0 in R. grylio and R. 

 catesbeiana; tympanum in males is proportionately greatest in R. grylio, 

 somewhat smaller in R. catesbeiana and R. clamitans and smallest in R. 

 heckscheri; intertympanic width of 95 mm. male in length (snout to vent) 

 4.52 in R. heckscheri, 5.43 in R. clamitans and 6.3 in R. grylio (R. catesbeiana 

 males of 95 mm. have tympanum poorly developed, but a 136 mm. has it 4.85) ; 

 in general, intertympanic width broadest in R. heckscheri and R. catesbeiana 

 and narrowest in R. grylio; distance from the rear corner of the eye to the 



