were long overlooked by earlier workers, and obvious specimens of 

 R. tricuspis were identified as R. lunulatus . 



Zumpt (l9i!.3A,1950A) accepted the differing descriptions of 

 R tricuspis and R. lunulatus as referring to separate species. 

 He synonySiz ed R. -glypbis under R. lunulatus but dxd not see 

 sieciB^ns of R rtrfeuspis . Theiler (^7) reared R. tricuspis 

 S rse^ed th at the gan ge of variation in the ^1 generation 

 from a single female included characters ascribed to R. tricuspis 

 arwell as to R. lunulatus . For the purposesof the present work, 

 Theiler has re-gt uilied her material and confirmed her earlier ob- 

 se?ia?Ions. Santos Dias (195QD,1952C) described differences, 

 which Theiler has found to be intraspecific, for R. tricuspis 

 and R. liinulatus . 



It might be indicated that in Theiler and Robinson (195^) 

 the term rT lunulatus is used only for literature references of 

 thit Sme- bul does n ot infer that these authors consider R. 

 lun\ilatus to be a valid species. 



In the following description, the range of variation in 

 Theiler's material is noted. Only typical specimens "ere des- 

 cried in her 19U7 paper, in which R. tricuspis was compared 

 with R. simas simus . 



IDENTIFICATION 



Male: The somewhat linear arrangement of the few ^^f' 

 deep -i^al puBctations identifies R. tricuspis m the R. simu^ 

 SoSp wSSn'Xch it staMs out easilF^^^curiously sinuous 

 gsterior margin of the adanal shields that are P-J-^^^ J^^^ 

 or less as a spur at the inner or outer juncture, or at both 

 "^IZeT. in'^pical specimens, these two PO^-f^J^JP^^.^ 

 the Dointed, heavily sclerotized accessory anal shields give 

 tSs^ea of the tick a tricuspid appearance. The spurlike outer 

 ^mctSe is pronounced but the inner ix>int is frequently more 

 r^S and shorter. Lateral grooves, usually ^^^l ^"^^?^^^ > ,^t 

 r^y may be much reduced or indicated by only a l^^^.^^^^^f ^f 

 ^ctations. The posterior grooves are typically distinct but in 

 C sp^:?^ns tenTto disappear. /"Note that in ^peci^- °^J- 

 simpsoS from a single host the same variation m scutal grooves 



_ 776 - 



