H.-M. WOODCOCK 15 



solid particles, it does so at any part of its bocly, more particularly in 

 the posterior région, in the same manner as does Cercomonas. There 

 are frequently prominent, deeply staining, rounded grains in the cyto- 

 plasm (cf. fig. 3), but thèse are unlike the Bacteria, etc., in the surrounding 

 niediam and do not, I consider, represent ingested food. It is only 

 necessary to compare the cytoplasm of, say, Monas, Bodo, or Phyllo- 

 mitus, with its contained Bacteria and other food-particles, to see the 

 différence. Thèse large, round granules, represent, rather, some product 

 of metabohc (probably nuciear) activity. Bat there can be no doubt, 

 I think, that this small accessory flagellum is comparable mth tliat of 

 paramastigote types. 



The nearest relationships of Helkesimastix are very difficult to déter- 

 mine. It resembles, in one character or another, several of thèse proto- 

 mastigine forms and yet difïers from them ail in some important respect. 

 On the whole, Helkesimastix is probably most nearly allied to Cercomonas, 

 although there is the important différence that it has lest the powerful, 

 long anterior flagellum of the latter type, but on the other hand has de- 

 veloped a short accessory flagellum, which Cercomonas does not possess. 



2. Alphamonas Alexeieff a synonym of Spiromonas Perty. 



Alexieff recently described^ a coprozoic Flagellate, which lie 

 regarded as an entirely new type and named Alphamonas coprocola, 

 n. g., n. sp. The author considered that tliis form had claims to be 

 considered intermediate (« forme de passage )>) between Bacteria and 

 Flagellâtes. 



In comparing Alphamonas with certain other coprozoic Flagellâtes, 

 Alexeiefï considers — and quite rightly — that his form is not related 

 to Helkesimastix. It is évident, however, that Alexeieff had not then 

 seen my later paper on coprozoic Flagellâtes (réf. above), or he would 

 certainly hâve realized that the form he describes is the same as that of 

 whose life-cycle I there gave a much fuller account, namely Spiromonas 

 angustata (Duj.)^. I made ail my original observations upon material 

 derived from sheep- and goat-dung ; but on noting that Alexeiefï obtained 

 his form from horse-dung, I started cultures of this material, and, as 

 expected, hâve had no difïiculty in obtaining Spiromonas. 



1. Arch. Zool. exp., vol. 57, 1918, N. et R. p. 1. 



'l. By an oversight, angustata is writtcn angunta iu my paper. 



