18 NOTES ET REVUE 



rentiation into nuclcar membrane, linin-reticiilum, and karyosome marks 

 a great advance on any type of nuclear structin-e found in Bacteria. 

 In the case of many nuclei of the protokaryon type, commonly found in 

 thèse simple Flagellâtes, practically ail the chromatin is contained in the 

 karyosome ; Spiromonas is no exception in this respect (cf. my figs. of 

 Bodo, Heteromita, Phyllomitus, etc.). 



As regards the life-cycle, I hâve shown above tliat there is i-eally 

 nothing comparable Avith (( epiplasm » in the multiplicative cyst of Spiro- 

 monas. The division into 3 or 4 daughter-individuals within a délicate 

 ectocyst cannot be correctlj^ compared with endosporous spore-formation 

 in certain Bacteria or Saccharomycetes. It is essentially an ectosporous 

 process-not the foimation of daughter-individuals or spores within the 

 body of the parent. It is much more comparable, for iixstance, Avith 

 the schizogony of Telospor dia (Ectospora)^ than with the spore-formation 

 in Neosporidia (Endospora). Lastly, conjugation (syngamy) occurs 

 in Spiromonas, as I hâve found to be the case in many other of thèse 

 coprozoïc Flagellâtes ; and up to the présent no évidence is forthcoming 

 of any such phase in the life-cycle of Bacteria. 



Spiromonas lias no cormection with Ancyromoiias. It is undoub- 

 tedly a Heteromastigine tjrpe, and best placed, I consider, in the Fam. 

 Bodonidae. Its nearest relationships are probably with Bodo and 

 Heteromita ; but it dififers from thèse forms in that the posterior flagellum 

 never appears to function as a passive trailing flagellum and salso in 

 regard to certain phases in the life-cycle. 



\. lu a^rtain Harmogregarines. schizogony takes place inside a délicate cyst-membrane. 



