46 WEST AUSTRALIAN PITCHEU-PLANT, 



Mesophyll. — Beneath the epidermis is a layer (4-5 cells thick) 

 of deeply-staining cells, containing much chlorophyll. This layer 

 contains few intercellular spaces, but there are small cavities 

 under each gland similar to those under each stoma. Beneath 

 this layer there is an open network of cells which contain few 

 chlorophyll granules, and stain lightly. This layer is similar to 

 that in the lid, and indeed in the middle layer of all parts of the 

 pitcher. Vascular bundles are very plentiful all through the 

 rim, but are most plentiful in the outer dense layer under the 

 epidermis, where also blind terminations occur, sometimes near 

 glands. After close examination of many preparations, however, I 

 am unable to say that they bear any constant relation to the 

 glands. In the loose interior tissue, wherever the vessels occur, 

 they are surrounded by a sheath of the dense tissue mentioned 

 above as lying under the epidermis. Starch is found plentifully 

 in these cells, and also, but less plentifully, in the middle layer. 

 The substance of the teeth, like that of the lid, and of the pitcher 

 body, is evidently actively assimilative. 



Epiclermis of collar. — This is of the same character as that on 

 interior of lid, but the conical hairs are longer, reaching their 

 greatest length at the interior angle where the collar joins the 

 interior surface. Each cell has a very distinct nucleus. There 

 are no glands, and yet the surface seems attractive to insects, as 

 they stay on it and lick it for a long time. 



Mesophyll oj collar. — The surface view just under the epidermis 

 shows regular hexagonal cells of large size, but none of the 

 granular deeply-staining cells of the corrugated roll. The walls 

 are thin, and the cells gradually pass into the loose network of 

 cells with little chlorophyll and a little starch. Forking vascular 

 bundles are found here, which terminate in loosely spiral blind 

 endings. Here also the vessels are invested with a sheath of 

 dense tissue. The mesophyll is so open that it can be seen to be 

 spongy with the naked eye. 



Interior of pitcher : upper glandular surface. — The whole of 

 the interior of the pitcher is very smooth and glossy, and as Prof. 



