30 bulletin: museum of comparative zoology. 



is not of such pattern as to point out these organs. It seems probable 

 that the nerves innervating these organs lose their sheaths before 

 reaching the epidermis. The exact innervation of these organs should 

 not be difficult to determine by one of the finer histological methods, 

 and oflPers an interesting problem. 



H. VISCERAL END ORGANS. 



The organs in which visceral nerves terminate, both efferent and 

 afferent, include glands and smooth muscle fibers for the former and 

 specific visceral sense organs and mucous membrane epithelium for the 

 latter. As no methods were employed to determine the actual nerve 

 terminations, a detailed description of these structures at this time 

 would have no significance for the general descriptive anatomy of the 

 cranial nerves which follows. In Anolis the mucous membrane of the 

 mouth cavity, the pharynx, and the nasopharynx presents a variety 

 of conditions in different regions which are demonstrable without the 

 use of special staining methods. It is throughout richly glandular, the 

 glands having the simple vesicular or tubular type along the gums and 

 the tongue, while in the postlingual region the epithelium is ciliated 

 and has a rich supply of unicellular glands of the goblet-cell type. 



The taste buds are confined almost entirely to the mouth cavity 

 proper, although an occasional bud was found in the region of the 

 larynx (Plate 4, Fig. 11, gm. gus.). They have the structure which is 

 typical for these organs elsewhere, possessing a well-defined gustatory 

 pit, in the base of which the sensory cells terminate. These taste 

 buds are distributed along areas which stand out as sensory -glandular 

 patches along the roof of the mouth and inner gums of both upper and 

 lower jaws (Plates 4, 5, Figs. 9-12, gm. gus. m., gm. gus. I.). Their 

 position is shown by the course of the sensory rami of nerve VII 

 (palatine and chorda tympani). No taste buds were found on the 

 tongue itself. Whether there are other sensory buds besides taste 

 buds, was a question that suggested itself through the presence of 

 clumps of cells which lacked the gustatory pit and were less sharply 

 defined but were not like glands in their structure. The fact of 

 importance in connection with visceral end organs is that typical 

 gustatory buds are readily demonstrated, and their distribution fully 

 determined in the series of sections from which the study was made. 

 Upon this are based certain conclusions as to the nature of some of 

 the branches of nerve VII. 



