46 bulletin: museum of comparative zoology. 



lively simple. The ganglion (Plate 5, fig. 15, gn. cil.) lies between the 

 membranous cranium on one side and the bursalis and retractor oculi 

 muscles on the other, and is separated from both III and V by roots 

 of considerable size. It is connected with nerve III, as is generally 

 the case in other forms, by a shorter, thicker root, radix brevis (Plate 2, 

 fig. 4, rx. cil. Ill), while the root from V {rx. cil. V) is longer and more 

 slender. Only the proximal end of this root is shown in the figure. 

 Both roots communicate with the ganglion directly and in like manner 

 the two ciliary ner^'es arise directly from its distal end. In the 

 plotting (Plate 2, fig. 4) this point is hidden by nerve III. These 

 ciliary nerves (Plate 5, fig^. 13_, 14, cil.) cross dorsad of the optic nerve 

 to enter the sclerotic coat of the eyeball. They keep close together 

 in their course as they pass laterad and cephalad around the eyeball 

 to their terminations in the striated muscles of the ciliary body and 

 epithelial surfaces. From the smaller of these ciliary nerves is given 

 off a very fine branch, which follows the others for a distance but is 

 lost before its entrance into the eyeball. In sections individual 

 medullated fillers are seen to be given off from this small ramus. 

 These are lost along the walls of the small blood vessels. The pres- 

 ence of these fine rami in immediate contact with the retractor oculi 

 muscle at its end of insertion could readily give rise to a misinter- 

 pretation in regard to their distribution (see p. 32). It is possible 

 that Osawa ('98, p. 537), basing his conclusions on dissections alone, 

 made such an error in his account of Hatteria. In Anolis every 

 recognized fiber leaving the ciliary nerves could be traced cephalad 

 beyond the most anterior extent of this muscle {rtr. oc), and in no case 

 w^ere these fibers distributed to the muscle. 



In many points the microscopic evidence was far from conclusive 

 for determining the relations of the ciliary roots and nerves to the 

 ganglion, but they offer certain facts worth recording. The short 

 root is a large one and is principally composed of fibers of very light 

 medullation, but not of the smallest caliber, i. c, they are larger than 

 those of the visceral sensory system as shown in palatine VII. Among 

 these are a very few coarser fibers of a medullation sufficiently heavier 

 to make them conspicuous. These are as large as many that remain 

 in the somatic motor rami, but do not equal in size th6^e which pre- 

 dominate in these motor nerves, nor are they segregated into a single 

 group to be readily followed through the ganglion. Howe^'er, the 

 fact that every section through the ganglion shows about the same 

 number of these coarser fibers points to the possibility of their unin- 

 terrupted passage. The ganglion is a uniformly oval structure_, the 

 cells of which entirelv surround the short root. 



