V 



54 bulletin: museum of comparative zoology. 



talis and the nasalis, Varanus agrees with Anohs in so far as no 

 branches are given off from the nasalis between the ciliaris and the 

 posterior extremity of the internal nares, where;, as she states, the " r. 

 comm. cum ramo platinus VII" is "composed of at least two distinct 

 fiber bundles," which communicate with palatine VII; then follow 

 the same divisions as noted for Anolis anterior to this, i. e., "r. latera- 

 lis" and "r. medialis," these having much the same distribution as in 

 Anolis. To the medialis are assigned the following branches, r. pre- 

 maxillaris superior (dorsal) and r. premaxillaris inferior, these inner- 

 vating the skin of the nose and the lips. 



Ethmoidal ganglion {gn. eth.). This is a ganglion of oval form and 

 one fourth mm. in length, which lies closely applied to the ventral 

 and lateral sides of the nasahs nerve (Plates 2, 3; Plate 4, fig. 10). 

 The ganglionic cells do not entirely surround this nerve, and there is 

 no passage of medullated fibers from nerve to ganglion. The gan- 

 glion is in connection with the median branch of the palatine nerve 

 through communicating bundles of fibers ; these fibers are also accom- 

 panied by sympathetic ganglion cells, which form a sort of column of 

 cells extending from the side of the ganglion opposite the nasalis 

 nerve to the palatine ramus (Plate 4, fig. 10). A dissection of this 

 ganglion and its connections, mounted in balsam, shows some fine, 

 lightly medullated fibers from the stalk passing both caudad and 

 cephalad in the palatine. The almost complete lack of medullated 

 fibers among the cells indicates that the ganglion has to do largely with 

 non-meduUated fibers. A very small bundle of such fibers joins the 

 ganglion on its posterior side close to the nasalis nerve. To what 

 extent this ganglion may be in connection with non-medullated fibers 

 of the nasalis, could not be determined, but such a relation would 

 appear to be the only explanation of the anatomical relations observed. 

 The cells of the ganglion are smaller and less clearly defined than 

 those of the cerebral ganglia, showing in this their sympathetic char- 

 acter (Carpenter, :06). 



The ethmoidal ganglion occurs regularly in birds, as far as they ha^'e 

 been studied, as a group of ganglion cells on the ophthalmic branch of 

 V. Bonsdorff ('52) gave it the name of "ganglion ethmoidale," and 

 Rochas ('85) "g. orbitonasale." In the goose Cords (:04, p. 59) de- 

 scribes this ganglion as being 1 mm. long and ^ to | mm. broad, and as 

 having the same connections as we find in Anolis, i.e., with the ophthal- 

 mic branch of V and the palatine branch of VII. There can be no 

 doubt of the homology of this ganglion as described for birds with the 

 structure to which the name has been given in zVnolis. 



