willard: cranial nerves of anolis carolinensis. 65 



(Plate 3, fig. 7), the plotting from sections (Plates 2, 3, figs. 4, 6) 

 being incomplete. Notwithstanding great variation in detail, the 

 scheme of the plexus seemed to be in the main the following : — 

 (1) Taking the main palatine as a point of departure, all the rami 

 have their origin from VII at one point, or nearly so. This is where 

 the palatine ganglion lies, as described by several authors. The 

 palatine here is a ramus of considerable size and immediately gives off 

 two main branches. These come off separately, but close together, 

 in the plotted series (Plate 2, 3, figs. 4, 6). In fig. 7, Plate 3, they 

 both arise from the ganglion. One of these {pal}) swings abruptly 

 outward crossing ventral to the infraorbital ramus of V (Fig. 7). It 

 innervates the mucous membrane and taste buds (gm. gus.) on the 

 way. The other (pair) keeps a more median position until it crosses 

 V to anastomose with the first {pal}) to form the lateral branch of the 

 palatine {pal. L). (2) There is an anastomosis {an'stm. pal. I.) with 

 V, from near this juncture, and sections always show ganglion 

 cells at this point (Fig. 4, cl. gn. sy.). (3) There are always two or 

 three branches (Fig. 7, a, 13, y) which turn back to join V just distal 

 to the infraorbital ganglion, and their appearance indicates a large 

 proportion of non-meduUated fibers. (4) Anterior to the orbit there 

 are anastomoses (a) that connect the intermediate branch of the 

 palatine with the maxillary nerve just before the latter passes into 

 the maxillary bone (Plate 2, 3, figs. 4, 6 and 7, an'stm. pal.i'm), and 

 (b) that connect the, median palatine ramus with ramus nasalis V 

 through the ethmoidal ganglion. These anastomoses account for the 

 mixed character of the lateral and intermediate rami of palatine VII. 

 Terminal twigs given off from the infraorbital plc.vus. Of more 

 importance than the exact form of the anastomoses are the twigs for 

 terminal distribution given off from the branches making up this 

 plexus. In all cases the main course of the principal branches is 

 explained by an examination of the mucous membrane of the roof of 

 the mouth. x\ll the rami carry viscero-sensory fibers for the innerva- 

 tion of the taste buds. These sense organs are distributed within two 

 restricted fields of the mucous membrane underlying the orbit, a 

 small median area and a lateral one. The median group of taste buds 

 (Plate 5, fig. 12, gm. gus. m.) is much more limited than the lateral 

 group. Here the buds are within a sensory-glandular patch little more 

 than a millimeter in length (between sections 732 and 874), which lies 

 on the pterygoid. The position is approximately indicated in the fig- 

 ure of the skull by the foramen in the pterygoid bone (Plate 1, fig. 3, 

 for. pt.). The innervation is from a twig {gm. gus.) of the median 



