68 bulletin: museum of comparative zoology. 



connection between the superficial sympathetic trunk of the head and 

 the infraorbital nerve. 



(2) Ramus hyomandihularis (hy-md.). This nerve leaves the genicu- 

 late ganglion (Plates 2, 3, figs. 4, 6, 7, g7i. VII) at its posterior end. 

 It embraces \'iscero-motor and viscero-sensory components. In cross 

 sections it is to be seen that these two components are entirely sepa- 

 rate throughout their course, the motor fibers having the more dorsal 

 position. The latter include all the motor elements of nerve VII; 

 these pass from the root across the dorsal side of the geniculate gan- 

 glion, not penetrating it, and then turn sharply into the ramus hyo- 

 mandihularis. The motor and sensory elements each take up about 

 half the area of a cross section of the nerve, the sensory components 

 being of course much finer in caliber than the others. The course of 

 this ramus is dorsad and caudad, following the same projecting ledge 

 of the prootic bone (Plate 1) as does the palatine, but in an opposite 

 direction (Plate 3, fig. 7) . This course leads it to a point just ventral 

 to the articulation of the quadrate with the prootic process of the 

 ear capsule (Plate 6, 7, figs. 19-20). Here occur the crossing and 

 anastomoses of the head sympathetic trunk {comn. ex.) with the hyo- 

 mandibular ramus of VII (Plate 3, fig. 7) . The superficial sympathetic 

 trunk {comn. ex.) from this point to the lachrymal plexus and infra- 

 orbital ganglion is called by Watkinson ( :06, p. 464) " ranms recurrens 

 nervi trigemini ad facialem " ; its continuation to nerve IX, the " ramus 

 communicans externus cum glossopharyngeo." 



In a series of sections of Anolis, in which fixation in this region was 

 excellent, it is shown with certainty that practically all the medul- 

 lated fibers in this sympathetic ramus, which in every way resemble 

 viscero-sensory fibers, pass the facial nerve in continuous course. 

 The facial, however, makes its way between the sympathetic fibers, 

 most of which cross the facial nerve mesally; a few only split away 

 from the others to cross it laterally, and then immediately rejoin the 

 main bundle. This is not a sympathetic center of any importance, 

 although from four to six very small ganglion cells are inclosed l^etween 

 the two nerves at the point of crossing. While there was no evidence 

 that any of the fibers in the part of the sympathetic ramus between 

 VII and V turned into any of the parts of VII at this point of union, 

 there does seem to be a strong indication that some of the fine medul- 

 lated fibers contained in the hyomandibular ramus of VII are continu- 

 ous with a part of those in the posterior section of this sympathetic 

 trunk, i. e., that between VII and IX. 



Attention is here called to the fact that cutaneous fibers are not 



