98 bulletin: museum of comparative zoology. 



those of the ventral roots of the spinal nerves, is further emphasized 

 in the third hypoglossal root and points to a very direct correlation 

 between the size of the fiber and its function. In this case there is no 

 doubt that the larger fibers run the shorter distance, thus contradicting 

 Schwalbe's ('82) law and confirming Dunn's ( :02, p. 323) results in the 

 spinal nerves of the frog. There may be, and probably are, other 

 factors entering into this particular case. Koch ('88) and Brandis 

 ('93) speak of a differentiation in the central nucleus of XII in the 

 medulla of birds, where a " dorsal" portion is separated from the rest; 

 and the suggestion is made by them that this may be correlated with 

 the syringeal musculature. In Anolis any relation of XII to central 

 nuclei bearing on peripheral differences must await a detailed study of 

 the brain. 



W. GENERAL SUM^vIARY. 



1. Anolis possesses the cranial nerves typical of the amniote verte- 

 brate with one exception; there was not discoverable any representa- 

 tive of the spinal accessory nerve described in other reptiles, and the 

 muscles innervated by this nerve in other forms seemed to be supplied 

 in Anolis wholly from spinal nerves posterior to the second cervical. 



2. The ganglia of cranial nerves V, YII, IX, and X are distinct 

 from one another and the roots of all issue from the cranium through 

 independent foramina. The ophthalmic ganglion also shows no 

 fusion with the other portion of the Gasserian ganglion. 



3. There is a wide distribution of sympathetic ganglion cells along 

 the afferent rami of the cranial nerves. These form definite ganglia 

 on palatine VII (palatine ganglion) on palatine VII and nasalis V 

 (ethmoidal ganglion), on maxillaris V (infraorbital ganglion), and on 

 mandibular V (mandibular ganglion). The topographical facts 

 would lead one to associate the development of these ganglia with 

 specialization of the glands of the head. No medullated nerve fibers 

 were found passing through the connective tissue surrounding these 

 glands. The presence of smooth muscle fibers in the head region 

 might also affect the development of the sympathetic. The sympa- 

 thetic system of the head in the matter of the arrangement of rami and 

 ganglia (as worked out incidentally to the study of the cranial nerves), 

 when compared with other described forms of reptiles, points to the 

 existence of a typical sauropsidan type of quite constant character. 



4. The nerve components (excepting the sympathetic) reach their 



