370 FERTILISATION OF THE GOODENIACE.^:, 



however, deeply coloured. The colour is very deep purple. The 

 indusium is two-lii3ped, and in mature flowers closes, except for a 

 small central opening. In all the older flowers I examined the 

 indusium was full of pollen, and the auricles lined with a sheet of 

 adhering grains. The indusium and stigma are purple, the style 

 below green. In no instance did I see the stigma outgrown, 

 or even so near the opening as to be capable of receiving pollen 

 from a visiting insect, so that this species also falls within the 

 group not fully adapted for insect-fertilisation; this is therefore 

 another species wuth arrangements for fertilisation complete 

 except at one point. 



Summing up, it appears that in Dampiera there is a complex 

 mechanism directed towards the accomplishment of cross-fertilisa- 

 tion by insects, and yet most species examined stop short of 

 completeness, from the stigma's not growing out so as to be exposed 

 to the touch of pollen-laden visitors. Those examined may be 

 divided into two groups; those having either a shallow indusium 

 so that the stigma may be reached by insects; and those in which 

 the indusium is deep, and from the stigma's not growing out, 

 incapable of being insect-fertilised. In the first of these groups 

 are D. linearis, sp. (?), eriocephala and juncea; in the latter D. 

 Brnwnii, striata, luteijiora, Linschotenii, loranthifolia, lanceolata, 

 and adpressa. 



The various species I have examined show a gradation in the 

 comj)leteness of adaptation of the various parts. Thus in the 

 auricles there is a progression from the simple fold with a slight 

 hollow in the centre {D. linearis) to the most complex arrange- 

 ment of folds, hairs and trichomes as in D. stricta and D. Brozvnii. 

 And in those species which show this gradual increase of adapta- 

 tion, there is also a regular augmentation of colour in the auricles, 

 from the simplest with a patch of faint colour in the centre, to 

 the deep purple spread all over the auricle in the most complex 

 forms. This is also the case in the style, which varies from green 

 to purple, and the indusium, from pale red to purple. The stigma 

 is always coloured, in which the genus cliifers from almost every 

 other member of the order. The indusium also varies from the 



