PAKAcOliOTOCA AKKHMAXl (waRUEN). 329 



epitlieliuni consists of exceedingly tall and slender cells, and 

 the inner portion of these appears to be gradually transformed 

 into a thick secretion which fills tlio lumen of the gland 

 (PI. XIX, fig. 64, s. ep.). 



The external male organs are shown semi-diagrammatically 

 in side view in PI, XIX, fig. 68. Here the oi-gans are partially 

 extruded. The chitinous penis or middle lobe [cii. p.) is seen 

 within a sheath (sh.) lined by a thin cuticle continuous with 

 the general exoskeleton. The lateral lobes have become 

 shifted in position, doubtless in association with the inverted 

 position of the abdomen ; the right lateral lobe (r. I. I.) is 

 almost ventral (uppermost) and the left lateral lobe {I. I. I.) 

 is nearly dorsal (figs. 66 and 67). 



When fully everted there is a ring-like swelling around the 

 base of the lateral lobes ; this is doubtless the "basal-piece " 

 of Sharp and Muir^ (PL XVI, figs. 11 and 12, b.p.). In 

 front of this, the soft integument of the abdomen bears a pair 

 of setose plates which we have already regarded as repre- 

 senting sternite X, while tergite X is perhaps modified into 

 the U-shaped gonapophyses at the base of the lateral lobes. 



In the mid-ventral line there is a conspicuous internal 

 chitinous rod (PI. XIX, fig. Q8, m. ch. r.), which is an ingrowth 

 from the venti"al recess of the genital vestibule. To this, 

 important muscles associated with the lateral lobes are 

 attached. The ejaculatory duct is very narrow and very 

 muscular. It enters the chitinous penis by a foramen situated 

 near the posterior end on the morphological ventral surface 

 (fig. 68). The ejaculatory duct runs through the relatively 

 wide chitinous penis, and near its apex it widens into a small 

 "internal sac '^ (Sharp) [i. s.) which is somewhat crumpled, but 

 by the action of certain muscle and probably also by fluid- 

 pressure the sac can be everted (PI. XVI, fig. 10, i. s.). This 

 internal sac is singularly small, and its reduced size and soft 

 condition are doubtless connected with the fact that there is 

 no definite bursa copulatrix in the female apparatus. 



' Sharp, David, and F. Muir, " Anatomy of the Male Genital Tube 

 o£ Coleoptera," ' Trans. Ent. Soc. Lond.," 1912-13, p. 481. 



