851 



THE POLLINATION OF GOODENIA CYCLOPTERA. 



(N.O. G00DENIACE.E.) 



By Archdkacon F. E. Haviland. 



Plate xciii. 



Goodenia cy dopier a R.Br., in the State of New South Wales, 

 is generally confined to the Western Plains. It is a decumbent 

 species, having axillary flowers furnished with indusia to the 

 styles, and with auricles to the upper lobes of the five-partite 

 corolla. These auricles are sufficiently induplicated to form a 

 pocket into which the indusium, at a certain stage, becomes in- 

 truded, as explained below. As appears to be the case in other 

 decumbent species, the indusium is inflected into the auricles, 

 whereas, in erect species, it appears that the indusium remains 

 erect, and the auricles press down over it. 



The pollination and fertilisation of certain species of Goodenia 

 have frequently engaged the attention of botanists, with occa- 

 sionally dissimilar results; and I may specially mention, among 

 the workers on the Australian flora in this connection, the labours 

 of the late E. Haviland, F.L.S., and Mr. A. G. Hamilton. Both 

 authors had treated of different species of Goodenia, and with 

 the following results, which are of interest to this paper. Mr. 

 E. Haviland, writing of G. ovata* in the year 1884, observes — 

 " The stigma covered by the indusium has re-entered [the corolla] 

 through the passage by which it had passed out, the division of 

 the upper lobe"; and, after referring to the ciliated indusium as 

 being intended to brush the pollen off an insect visiting the 

 flower, came to the conclusion that the flowers are cross-fertilised. 

 I compare this with the observations noted by Mr. A. G. Hamil- 

 ton! in the year following the above, and who says, of G. heder- 

 acea^ that " the basal portion of the style is bent upwards so as 



* These Proceedings, ix., 449. 

 fOp. ciL, X., 157. 



