190 PROCEEDINGS OF THE ACADEMY OF [1899. 



long, terminating in a very short jiagellum and the long free vas 

 deferens. 



9 system : Vagina moderately long, with internal folds continu- 

 ous with those of the atrium. Spermatheca long, not differentiated 

 into duct and sjyerviatheca proper, but consisting of a long, some- 

 what folded tube of nearly equal diameter throughout, the apex 

 free and lying near the heart. Talon simple, vermiform. Other 

 organs as usual (fig. 1). Measurements of the principal organs, 

 pulled straight, are as follows: Length of penis, 10^ mm. ; of 

 epij)hallus, 31 ; of flagellum, 2 mm. Length of spermatheca, 27 

 mm., including the inseparable duct. 



The lung is of normal proportions ; there is a single large pulmo- 

 nary vein with numerous much smaller branches on each side ; 

 heart at base of kidney, the pericardium thin and transparent. 

 Kidney small and rather short, though slightly over double the 

 length of the heart, with distinct reflexed ureter, the " secondary 

 ureter" (' ^ Darmharjileiter " ) apparently closed throughout. 



Jaw (fig. 2) quite arcuate, with about eight ribs denticulating 

 the basal margin, the ends free from ribs. 



Radula with 26-1-26 teeth (fig. 3), the rhachidian row tricuspid, 

 the side cusps small, laterals with the usual large mesocone, small 

 ectocone and rudimentary entocone, marginals with the mesocone 

 bifid at tipj the outer ones with the ectocone split. The transition 

 from laterals to marginals occurs from the eleventh to the thirteenth 

 teeth. 



, Affinities of the Genus.^ 



The appearance of the shell is so completely that of Polygyra 

 that in attempting to decide its aflSnities one approaches the ques- 

 tion somewhat prejudiced. Turning to the genitalia, we find that 

 the male system is completely that of Epiphragmophora and the 

 Belogonous Helices generally. The well-developed epiphallus and 

 the short but unmistakable flagellum are structures which are wholly 

 foreign to Polygyra.* Again, the long spermatheca with its apex 

 lodged near the heart, utterly unlike the short and difierently 

 situated spermatheca of Polygyra, is like Epiphragmophora and 



» By H. A. Pilsbry alone. 



* The genitalia of nearly every United States species of Polygyra are 

 known, either by the published work of Mr. W. G. Binney or by MS. 

 descriptions and drawings of Messrs. H. A. Pilsbry and E. G. Vanatta. 



