EINAR LÖNNBERG, AN ATOM Y OF THE RUMINANTS. 5 



than to tlie »ovine» type, in contrast to Cephalophince espe- 

 cially Cephalophus. Although this is less pronounced in 

 Ourebia (conf. above) this genus as well differs a great deal 

 from tlie »ovine» type through the absence of the cushion at the 

 penis end and through the shortness of the processus ttrethra- 

 lis. It might thus be supposed that all members of the sub- 

 family Neotragince have reached approximately the same 

 stage of development and in this respect are more advanced 

 than Cephalophus 



In the subfamily C ervicapriiioe the male organ of Cobus 

 defassa is known as it has been described and figured by the 

 present author (6). The same organ of Cobus ellipsiprymnus 

 from the Zululand is, as might be expected, very similar. The 

 terminal portion has a well developed cushion vvhich bends 

 down at the tip and forces the urethra, which so far has 

 run mesially on the lower side of the penis, towards the 

 left side where it ascends and simultaneously detaches itself. 

 The free urethral prolongation thus measures about 33 mm. 

 Basally this processus urethralis measures fully 6 mm. in 

 thickness but tapers gradually to a httle more than 1 mm. 

 at the tip. From the thick base runs on either side of the 

 urethra a recurrent band which extends along the penis itself. 

 The band of the left side is thicker than that of the right 

 which is quite thin and also shorter than the other. As this 

 shape of the penis has been described from two different 

 species of the genus Cobus it must be assumed to be the 

 regular occurrence in the Water-bucks, but it is not a cha- 

 racteristic of the whole subfamily as will be proved in the 

 following. 



The penis of the »Reedbuck» {Redunca arundinum Bodd.) 

 is not very similar to that of Cobus. It appears to be more 

 similar to that of Sylvicapra grimmia. The terminal cushion 

 is not strongly developed. It is not thicker than the proxi- 

 mal portion of the penis, and it is not bent down at the 

 tip so as to force the urethra abruptly to the side as in 

 Cobus. On the contrary the urethra ascends gradually from 

 its mesial position of the under side on the left side of the 

 penis-end where it is received in a groove. By this arrange- 

 ment it is not much conspicuous that about 4 mm. of the 

 urethra is detached from the penis forming a processats ure- 



