CARBONIFEROUS LYCOPODS AND SPHENOPHYLLS. 



63 



(LepidophyllamT) major, Brongt. Fig. 9. In this latter species I 

 believe are sometimes included leaves of 

 Lepidophloios as well as bracts. 



In Lepidostrobus, as generally em- 

 ployed, are almost certainly included, 

 not only cones of Lepidodendron, but 

 also of Lepidophloios and Bothrodendron, 

 if not even of Sigillaria, and I strongly 

 suspect that the cones with naked pedicels 

 figured by me from the Ayrshire Coal- 

 field under the name of Lepidostrobus 

 (?) spinosus may belong to Sigillaria, 1 but 

 without some knowledge of the structure 

 and arrangement of the sporangia it is 

 impossible to determine their true sys- 

 tematic position. 



When perfect specimens of Lepidos- 

 trobi are examined the lower portion of 

 the cone contains the macrosporangia and 

 the upper the microsporangia, 2 the macro- 

 spores representing the female and the 

 microspores the male elements. It is of 

 course quite possible that some cones 

 produced only microsporangia and others 

 macrosporangia, and one whose structure 

 was described by the late Prof. Williamson 

 is supposed to have been of this nature, Fig. 9. — Lepidophyllwn 

 but the supposition may rest on imperfect ( Lepidostrobu, 

 knowledge. 3 Bron S t - 



When the bracts are removed from the 

 axis it exhibits a number of close, spirally-placed, small, elongated 



1 Trans. Hoy. Soc. Edit., Vol. XXXVII., p. 341, PL II., fig. 7 ; PL HI-, 

 figs. 11-12. 



2 See Williamson, "Organization of the Fossil Plants of the Coal 

 Measures," Part xix. Phil. Trans., J 893, Bot., PL VIII., figs. 51-52 = 

 cone of Lepidodendron orevifoiium, Will, (not Ett. ) ; also Binney, Palgeont. 

 Soc., Observations on the Structure of Fossil Plants, Part ii. PL IX., 

 fig. \ = Lepidostrobus Russellianus, Binney. PL X., fig. \ = Lepidostrobus 

 levidensis, Binney. PL XI., fig. 2= Lepidostrobus Wunschianus, Binney. 



" Maslen, "The Structure of Lepidostrobus,'' Trans. Linn. Soc. London, 2nd 

 Ser. Bot., Vol. V., Pairt xi., p. 358. 1899. {Lepidostrobus Oldhamius, Will.) 



(No. 



Natural size. 



2527.) 



