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IV. — On Evidence of a New Species of Titanosuchus (T. cloetei). 

 —By E. Broom, M.D., B.Sc, C.M.Z.S. 



In the collection of the South African Museum is the front portion 

 of the left mandible of a large Theriodont reptile. It was presented 

 to the Museum many years ago, by Mr. Justice Cloete, and the 

 locality at which it was obtained is noted as the " Gamka Eiver." 

 The specimen has been so beautifully weathered that though all 

 the matrix is gone the bone is still in perfect preservation. The 

 remains of four incisors, the canine, and four molars are preserved, 

 though unfortunately the crowns of all the teeth are lost. The jaw 

 and teeth agree in their general features so closely with those of the 

 type of Titanosuchus ferox that there is little doubt the two specimens 

 belong to the same genus, though the difference in size is such that 

 the specimen most probably belongs to a new species, which I 

 propose to call Titanosuchus cloetei. Owen, with his usual thorough- 

 ness, gives detailed measurements of the larger teeth in T. ferox, 

 and the difference in the two species can be best appreciated by 

 contrasting the dental measurements. 



T. ferox. T. cloetei. 



1st incisor Imperfect. 21mm. x 14 mm. 



Interval between i. 1 & i. 2 10 mm. 10 mm. 



2nd incisor 21mm. x 15 mm. 18 mm. x 13'5 mm. 



Interval between i. 2 & i. 3 10 mm. 11 mm. 



3rd incisor 20 mm. x 12 mm. 20 mm. x 14 mm. 



Interval between i. 3 & i. 4 18 mm. 8 mm. 



4th incisor 23 mm. x 17 mm. 18 mm. x 14 mm. 



Interval between i. 4 & c. Not given. 9 mm. 



Canine 50 mm. x 35 mm. 30 mm. x 26 mm. 



In T. cloetei the four molars occupy a space of 40 mm., and each 

 root is practically round and measures about 8 mm. in diameter. 



From the above measurements it will be seen that the incisors in 

 T. cloetei are appreciably smaller, and that the canine is much 



