AND PECULIAR FORM OF THE EMBRYO IN THE CLUSIACEE. 249 
I have already alluded to the existence of two somewhat different developments of the 
seed in the Clusiacee; the foregoing description affords an example of the one which 
includes all cases (as in the tribe Clusiew) where a number of seeds are formed in each 
. cell of the ovary, and where they are attached in a horizontal position by their base to 
the axile placenta. The other development occurs in those instances (as in the tribes 
Tovomitee and Garcinieæ) where only one seed is formed in each cell, and where this is 
fixed to the axile column in a vertical position by its ventralface. For reasons that will 
be offered in another place, I propose to exclude the Moronobeæ and Calophyilee from 
the Order. In the first case (the Clusiec), the raphe, enclosed within the fleshy arillus, 
is seen to extend from the base to the apex of the seed, and is free from the testa; in the 
second instance the testa is thinner and more membranaceous in texture, and enveloped 
in an overlapping arillus, which is thicker and more membranaceous in substance; it has 
a large hilum upon its ventral face; the raphe, less discernible, is imbedded in the sub- 
_ Stance of the testa, where it spreads into numerous branching nervures, conspicuously 
extending over its surface: in the Garciniee we find a similar testa, enveloped in an 
entire, copious, more or less mucilaginous pulp; tolerably good figures of this develop- 
ment are given by Gærtner in plate 105, illustrating the seeds of Garcinia, and in several 
plates of Dr. Wight’s ‘Icones.’ Were it not for the explanation afforded by the analysis 
of the Clusiec, the structure of the embryo in the other tribes, Tovomitee and Gareiniee, 
would not be so easily understood. : During my residence in Brazil, I examined in a living 
state the fruit and seeds of different species of Zovomita, Commirhea, and more especially 
of a plant which I have called Lamprophyllum letum, the type of a genus very distinct from 
. Garcinia, and comprising numerous species of South American and West Indian origin, 
among them the Calophyllum Calaba of Linnsus, and others associated with Gareinia 
and Calophyllum, which last genus I consider to be foreign to the Order. The analysis of 
the seed of Lamprophyllum will afford a good example of the second mode of development 
above referred to. "The drupe is here about the size of that of Calophyllum Calaba, as 
figured by J. acquin (Stirp. Amer. tab. 165), and contains generally two, or often by abortion 
a single seed, about the size of the kernel of a hazel-nut, which is enveloped in a thiek 
mucilaginous pulpy arillus: the testa is thin and brittle, marked by numerous veins 
branching from the ventral hilum, and it contains a solid nucleus of a firm and somewhat 
fleshy consistence, exhibiting in the apex a minute prominent nipple of the size of a very 
small pin's head, seated in a deep hollow depression, a little below the summit towards 
the ventral face; near the base, somewhat on the dorsal side, is seen another smaller 
speck, which is green and shining, exactly like that described in the nucleus of the 
Clusiee. On making a longitudinal section, the main body of the nucleus is seen to be of 
a pale sulphur colour, studded with numerous small oblong ducts, whieh, when thus cut, 
copiously exude a deep yellow viscous juice: a slender terete neorhiza, exactly resembling 
because we see in the Clusiee that the neorhiza is traceable only to the nascent point of the plumule, that the eotyledons 
are wholly exserted from and an extension of the main body, and that many of the striæ or Jong tubular ducts, which 
extend from the base to the apex of the mass of the nucleus, are carried VE interruption along “ie outer face of 
the cotyledons, proving the continuity of the one with the other; for were it of the nature of albumen, it would be in 
the form of an investiture of the embryo, not a prolongation of it. 
