^S-1 



THE CILLATES 



a flat or slightly concave right side. This 

 species has 9 formae which differ prin- 

 cipally in their caudal spination: E. ecaiid- 

 atiuii forma ecaiidattiiu has no caudal 

 spines, forma caiidalioii has a single long 

 spine and the posterior end of its body is 

 narrow, liama/io)/ has a single long spine 

 and the posterior end of its body is broad, 

 bulhiferiim has a bulb-shaped appendage 

 instead of a spine, bicaiidaln))i has 2 

 spines, IricaudalKiii has 3, quadricauda- 

 tum has 4, catteueoi has 5 short ones, and 

 fasciculus has 5 very long ones with 

 greatly swollen bases. All 9 formae occur 

 in the rumen of cattle, the most common 

 being ecaudatu)ii and caudalu))/. Ecauda- 

 tum, lianiatum and cattaiieoi are found in 

 sheep, and the last in goats as well. 

 EcaudaluDi and caudatuni also occur in the 

 reindeer, caudatu>ii and luiDtaluni in cam- 

 els and caudatum in the elk. 



Genus EODINIUM 

 Kofoid and Maclennan, 1932 



The metoral zone of membranelles is 

 at the same level as the adoral zone. 

 Skeletal plates are absent. The macro- 

 nucleus is a straight, rod-like body near 

 the left edge. Two contractile vacuoles 

 are present. This genus occurs in the 

 rumen and reticulum of cattle and sheep. 



E. posterovesiculatu))i (Dogiel, 1927) 

 Kofoid and MacLennan, 1932 measures 

 47 to 60 by 23 to 30 pt and has a relatively 

 long, flattened body with rounded ends. 

 The macronucleus is very long, straight, 

 and has 2 deep depressions on its left 

 side. The micronucleus lies in the pos- 

 terior one, and a contractile vacuole in 

 the anterior one. The second contractile 

 vacuole is posterior to the macronucleus. 

 This species occurs in cattle. 



E. bilobosum (Dogiel, 1927) Kofoid 

 and MacLennan, 1932 measures 46 to 60 

 by 30 to 44)ji and has a relatively short, 

 flattened body with 2 caudal spines, 1 

 dorsal and the other ventral. The nuclei 

 and vacuoles are similar to those of E. 

 posterovesiculalum . This species occurs 

 in cattle and sheep. 



Eodiniuiu lobaluiii Kofoid and Mac- 

 Lennan, 1932 measures 44 to 60 by 29 to 

 37 (i and has a narrow body. The macro- 

 nucleus is narrow and rod-like, and is 

 almost as long as the body. It has 3 large 

 depressions in its left side; the micronu- 

 cleus lies in the middle one, and the con- 

 tractile vacuoles in the end ones. This 

 species occurs in the zebu. 



Genus DIPLODINIUM Schuberg, 1888 



The metoral zone of membranelles is 

 on the same level as the adoral zone. 

 Skeletal plates are absent. The macro- 

 nucleus is beneath the upper surface of the 

 body; its anterior third is bent to the right 

 at an angle of 30 to 90". Two contractile 

 vacuoles are present. This genus occurs 

 in the rumen and reticulum of cattle, goats, 

 sheep, camels, reindeer and various wild 

 ruminants. 



In this genus, as in Entodiniuni, 

 there is also considerable variation in the 

 caudal spination. 



DiplodiniuDi dentatuiii (Stein, 1858) 

 Schuberg, 1888 measures 55 to 82 by 44 

 to 62 ji . Its body has a broad, truncate 

 posterior end with 6 large, relatively 

 heavy, incurved caudal spines. The left 

 side is convex and the right one concave. 

 The macronucleus is 25 to 50 (Lt long; it is 

 heavy and rod-like, with the anterior end 

 bent at a 45° angle. The 2 contractile 

 vacuoles lie in the left rib slightly below 

 the midline. This species occurs in the 

 ox and zebu. 



D. quinquecaudatum Dogiel, 1925 

 measures 57 to 73 by 47 to 65/i . It re- 

 sembles D. deu/atu)!!, but has 5 caudal 

 spines. It occurs in cattle and sheep. 



D. anacaiilhuDi Dogiel, 1927 meas- 

 ures 60 to 124 by 38 to 72 /i. The poster- 

 ior end of its body tapers, giving it a 

 somewhat conical appearance. The ma- 

 cronucleus varies a good deal in length. 

 Its anterior third is bent at an angle of 

 about 45°. The 2 contractile vacuoles 

 are on the lower side. This species has 

 7 formae (which Kofoid and MacLennan, 



