238 REMARKS ON THE FERTILIZATION OP THE GENUS GOODENA, 



paper, I speak of the upper lobe, and I point out, that, while the 

 lower lobe is not divided so deeply, the upper one is divided nearly 

 to its base. This of course makes the claws of the lobe very narrow 

 and weak, so that the edges of the two divisions which only touch 

 each other, ai'e, as I said, liable to be separated by the slightest 

 movement. Generally however, this opening of the divisions of 

 the lobe is not necessary, the indusium and stigma being in most 

 cases already within the corolla. I only say in my paper •' some- 

 times it happens, that although the mouth of the indusium is 

 presented to the opening of the corolla lobe, it may not have quite 

 entered it, and that, in such case, the ready opening of the divisions 

 of the upper lobe remedies the evil, and exposes the stigma to the 

 visit.s of insects from the inside of the corolla" ; but this (happening 

 so exceptionally) is a matter of little importance. A. more material 

 point is that which Mr. Hamilton mentions as a "significant fact" 

 that neither full anthers or empty indusia are ever found in open 

 flowers. On reference to my paper, above referred to, it will be 

 found that I say " Taking now a mature d^nd fully expanded jiower, 

 weshall find the stamens still outside, and usually bent farther away 

 from the corolla, the anthers all open and, and the pollen either in 

 abundance, or, in some cases, past that stage, all gone, and the 

 anthers shrivelled." Of course I intended by this to imply that 

 the pollen was generally exposed in the open anthers, but that in 

 some cases insects had carried it away and left the anthers empty. 

 As to the non existence of empty indusia in open flowers. I find 

 in my rough notes, made while studying the genus, a remark 

 touching an open flower of G. ovata thus " Notwithstanding that 

 the anthers are empty, there is not a particle of pollen either on 

 or inside the indusium or on the stigma ; showing that the flower 

 has not been fertilised by its own pollen (for the stigma was not 

 mature) and moreover that it has not yet received pollen from any 

 other flower." I found many flowers in this state, but I did not 

 deem it necessary to make a note of more than one. 



Perhaps the best authority on the fertilization of jjlants was the 

 late Herman Mueller, and I am bound to admit that in his work 

 on the fertilization of flowers he agrees with IMr. Hamilton, in 



