1903.] NATURAL SCIENCES OF PHILADELPHIA. 391 



in numerous oblique rows upon the surfaces of the coronal lobes. In 

 Floscularia there is in most species a single row of cilia upon the 

 lobes or on both the lobes and the interlobular coronal margin; 

 in a few there are two rows upon the coronal margin {pe- 

 lagica Rousselet, hoodii Hudson, cucullata Hood, trilohata Collins). 

 All these cilia are preoral; they constitute, therefore, a trochus, and 

 there is no postoral row (cingulum) on the external surface of the body 

 behind the mouth. In fact, no postoral cihary ring occurs, for the 

 diaphragm of Stephanoceros and Floscularia, wliich bears a row of 

 cilia, and on that account was homologized by Hudson and Gosse 

 with the cingulum of other Rotatoria, belongs to the intestinal tract 

 and not to the coronal margin at all. In the young of Apsilus and 

 Floscularia the mouth is ventral instead of terminal, as in the adult, 

 and the anterior region of the ahmentary tract is cihated, these cilia 

 not persisting to the adult stage. The diaphragm separates in 

 Stephanoceros and Floscularia an anterior infundibulum from a poste- 

 rior vestibulum, both without cilia; in Apsilus there is no diaphragm 

 and no distinction of two chambers, and also no ciliation. In two 

 species of Apsilus, in Floscularia and Stephanoceros an oesophageal 

 tube is present. On the vestibulmn follows a non-ciliated proven- 

 triculus, the posterior end of which is specialized as the mastax; next, 

 a ciliated stomach; then a non-ciliated posterior intestine, then the 

 short rectum opening into the cloaca. The intestinal ciliation is thus 

 limited to the diaphragm, the stomach and (in some species of Flos- 

 cularia) to the rectum. 



The foot is well developed in all but Apsilus; in this genus it is larger 

 in the young than in the adult, so has probably degenerated. Only in 

 F. chimccra Hudson does the foot terminate in two toes (it is doubtful 

 whether this form belongs in the family). In all other forms it termi- 

 nates in a single peduncle (except in Floscularia atrochoides Wier- 

 zejski). The foot is strongly contractile, but not retractile into the 

 trunk. The nephridial system is essentially alike in all three genera, 

 likewise the nervous system; no ring nerves have been found in the 

 coronal margin, and there is no suboesophageal ganglion. A dorsal 

 and a pair of lateral coronal sense-organs are always present, and an 

 additional pair of lateral ones in Apsilus. The germarium and oviduct 

 are unpaired; the germarium is small and rounded in most species, but 

 much elongated in Apsilus hucinedax (Forbes) and Floscularia ambigua 

 Hudson. Distinct foot glands are absent in the adult; the only glands 

 connected with the alimentary tract are one pair of stomach glands. 

 A gland around the dorsal sense-organ is found in some forms. 



