36 DR. J. D. HOOKER ON THE GENERA AND SPECIES 
much in these as in the female flowers, but though they never attain the same develop- ; 
ment as they do in the males, they approach it, and often contain vascular cords. In. 
one hermaphrodite flower (fig. 9) I found six perigonial scales, symmetrically disposed 
round the summit of the ovary. 4 
There is thus in a complete flower of Oynomorium a superior perigonium, an epigynous . 
stamen, a pistil consisting of one carpel with one simple style, and one pendulous ovule, - 
succeeded by an albuminous seed :—characters common also to Hippuris, and indicating | 
an affinity I have elsewhere noticed, and endeavoured to support by the structure of other 1 
Balanophoree. y ; 1 
I may here repeat what I have stated in the * Flora of the Canary Islands' concerning 
the pistil of Cynomorium; namely, that the simple concave style with two parallel vas- 
cular cords terminating an ovary which undoubtedly consists of one carpellary leaf, is à. 
strong evidence of the compound nature of a style of the simplest type; and that the two 1 
lateral stigmata are here perfectly obvious: which agrees with Mr. Brown's remarks on 
the composition, &c. of the pistil* (Plant. Jav. Rar. p. 110 in note). The stigmatic tissue - 
runs down the mesial line of the style, occupying the canal, and is covered by a very - 
delicate epidermis. 4 
I have never succeeded in tracing the development of the ovule in Oynomorium. The 
structure of the ripe seed has been determined by Richard, Lindley, and Weddell; but | 
admits of some little further illustration. In the first place, the embryo when fully ripe 
is considerably larger than is figured by any of these authors, and is never exactl 
globular, but sometimes broadly conical, the narrow end being placed next to the firm 
cellular integument of the seed. It consists of large cells with dark nuclei, full of o 
and presents no integuments whatever: it lies in a cavity of the albumen, nearer the base 
albumen as mucilaginous, and the cells 
either microscopically, or by the iodine test, whi en applied to the most delicat 
slices of fully formed albumen, turned its granular cell-contents brown. Mr. Weddell 
states that the albumen as well as the embryo contains oil (Ann. Se. Nat. 7. c. 178) 
“dans les Balanophora, dans le Oynomorium, et dans le Sarcophyte, j'ai rencontré un 
albumen charneux ou huileux, et un embryon de méme nature.” — 
Å On showing Mr. Brown my analysis of the style of Cynomorium, he informed me that this was a case he had had 
m view, and that he considered it strongly confirmatory of his theory. 
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