IN THE CLUSIACEÆ, MAGNOLIACEÆ, ETC. 87 
view of the question is one which certainly ought to have its due importance, and is one 
which I have never doubted in the smallest degree, viz. that Dr. Gray has watched the 
progress of the growth of the seed from the ovule and could never detect any subsequent 
‚or extraneous production over the primine. In the fullest credence of this assertion I 
still think we have convincing proof that such extraneous growth, though he failed in 
distinguishing it, must have taken place imperceptibly, in the same way that other 
observers have witnessed a similar gradual extension over the primine. 
The foregoing discussion has induced me to offer a few additional observations on the 
nature of the raphe. I have been led into this inquiry by the phænomena that present 
themselves in the Clusiaceæ, where I found it difficult to assign a reason, why the raphe 
should in one case consist of à single cord of vessels, almost in a free state, within the 
inner pellicle of the external arilliform coating, and wholly free from the testa, as inva- 
riably occurs in the C/usieæ, while in the Tovomiteæ, the raphe has no connexion with 
the enveloping aril, but appears imbedded in the slender chartaceous testa, not as in the 
former case like a simple cord, but spread in the form of several branching nervures con- 
tinually subdividing themselves, and distributed in new ramifications throughout the 
whole area of the testa. There appears here a difference of structure somewhat inexpli- 
cable, but after a little consideration, we may trace to one uniform rule the varying 
appearances of the raphe under these different forms and position. We owe to the pro- 
found investigations of our great countryman Mr. Brown, most of the knowledge we 
possess regarding the growth and development of the ovule and its original tunies, the 
changes they undergo during and after the anatropal metamorphosis, the gradual trans- 
formation of these tunics into the different coatings of the seed, and the share they afford 
in the nourishment and development of the growing embryo: but the phenomena 
attendant on the formation of the raphe, the modifications which these nourishing vessels 
undergo, and the different modes of their distribution, appear to have excited little notice, 
for we find few observations on the subject in the works of physiological botanista, oe 
this account I will venture to offer some remarks that have occurred to me in my pend- 
ing investigation of the Clusiacee. 
Let us now examine some of the circumstances attendant on 
Prior to this action, the body of the ovule or nucleus is generally ee ir aii 
an as d secundine are given : these cups gra 
are Am m co and conceal the nucleus, also called 
Size, and grow into complete tunies which envelope : . 
the ing: during iis ende stage, the vessels destined to nourish po Er hagen me 
diately from the placenta into the gangylode or common po 2 cues, nl 
With the secundine and primine then in contact with the ucl n iei 
gress of inversion, the gangylode, as before — zum "ue i e side of the 
‘and a portion of the placenta is drawn away with it and extended ove een 
primine, at first in the form of an elongated sheath or not d hide in the 
Carrying with it the spiral vessels proceeding from the — A quise cuiu fe its 
now remote gangylode: this placentary extension, whatever TTT at it may 
subsequent growth, must always remain outside the primine no ee de ia 
me agglutinated to it. Now these nourishing vessels may 
the inversion of the ovule. 
