[ 4E ] 
XXXIV. Notes on the Structure and Affinities of Batidex, Callitrichaceæ, Vorne. 
| and Cassytheæ. By B. CLARKE, Esq., F.L.S. 
Read June 16th, 1859. 
1. On the Structure and Affinities of Batideæ and Callitrichaceæ. 
THE principal points in the structure of the genus Batis, P. Br., have become well known, 
but its affinities have not been decided, while those which have been suggested for it are 
proved to be, for the most part, if not entirely, impracticable; further analysis therefore 
seems required before its affinities can be brought to light; and the following additional 
particulars, as regards its structure, and especially the relative position of the ovules to 
the axis of the ovary, will, I believe, make it evident that it must be a near ally of the 
Verbenacee. 
The male flowers are arranged in 4-rowed spikes, giving them a habit like that of the 
Verbenacee with 4-rowed spikes of flowers; but in one species the inflorescence becomes 
compound, which gives it in some degree the habit of a Globularia, as of G. orientalis. 
_ The calyx is reduced to one sepal, posterior ; but, since in the Acanthaceæ also the calyx is 
sometimes very nearly obsolete, this negative character is of little value. The corolla is 
attached by a very slender basis to the calyx, and consists of four petals with long claws 
. and oval or almost circular laminæ, and of four stamens alternate with them, two of 
which are placed anterior and posterior, and two right and left of the axis *,—the two 
lateral stamens having the rudiment of an ovary slightly adhering to them, which is 
sometimes bifid, the segments being lateral. 
The female flowers gti siterip which are 4-rowed, as in the male. The floral 
envelopes are deficient; but each ovary has at its base a quickly deciduous bractea rar 
LXVIII. fig. 27), so that in the early stage the spike is completely enveloped in — 
and is also terminated with three or four pairs of minute flowerless scales. | 'The rare 
are two, right and left of the axis, and the cells of the ovary four, so placed aon idi 
them stand right, and two left of the axis, so as to correspond in their deniqne T ot 
two stigmas, each pair having no bundle of vessels between them, but m vhi 
by very distinct bundles from the opposite pair (fig. 27 a). Each pek LÉ kii 
anatropal ovule, the raphe and foramen of which are very distinct. : e wn ER 
in relation to the inner angle of each cell; but in each pair of cells the rap 
: Labia ving beyond 
turned towards each other (fig. 27 a), as in the Ver nn rar ped ae ls y 
i , consis is sona MD 
doubt that the structure of the ovary is the same, co cin Vostoitües vily tà being 
Accordi is view of its structure, Batis differs from | 
di rense tole unless it is in the position of the stamens; for in the female 
* 7 h l 1 s LPG $ og> Le 3 - ' 
3 > 1r posi 10n in B. mar ı > y y q * 
diclinous and polypetalous, 
