p. K. ANOKHIN 



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reticular formation (Magoun, 1952; Moruzzi, 1956, ct al.), the local 

 limb lifting, i.e. the local conditioned motor act is cftcctcd by the 

 pyramidal tract and has a definite cortical localization. 



The question is: how do both these organic components integrate in the 

 animal's integral conditioned motor act? We endeavoured to answer this 

 question by means of a number of variations of conditioned reflex 

 experiments. 



In order to trace the very fact of the shift in the centre of the body's 

 gravity during the stage of formation of tlie positional excitation we 



Fig. 13 

 Stand designed with four separately recorded stages. In 

 addition to recording the spread of positiiinal excitation this 

 stand permits of recording the secretion of sahva and the 

 vegetative components. 



designed a stand by means of which we could record the pressure exerted 

 by each of the four limbs (Fig. 13). The experiment showed that the 

 moment the conditioned stmiulus was applied there was a rapid and 

 rather complex redistribution of muscular efforts, which could be easily 

 recorded kymographically and elcctromyographically (Koryakin, 1958). 

 It also showed that, if the conditioned motor defensive act was effected, 

 say, by the right hinci limb, the lifting of the paw was always preceded by 

 an uncommonly stable complex of relations in the excitations of the 

 brain stem. Now, if both sensomotor areas of the cerebral cortex are 

 removed, an interesting dissociation between the positional and local 



