NUCLEI AND KINETIC ELEMENTS 



99 



peared {Trichomonas, Kofoid). The term is certainly justified ia 

 connection with the remarkable kinetic structures of flagellates 

 belonging to the family Trichonymphidff'. In Trichonympha cam- 

 panula, Kofoid and Swezy (1919) describe the system as composed 

 of an external coating of cilia-like motile organs, three zones of flagella 

 with their basal bodies, rhizoplasts connecting basal bodies with a 

 great anteriorly placed blejiharoplast, and more deeply-lying myo- 

 nemes which apparently are not connected with the blepharoplast 

 (Fig. 50). Kofoid and Swezy regard the central organoid as a kind 

 of superblepharoplast, calling it the "centroblepharoplast" since it 

 has the attributes of a centrosome. When it divides the entire 





Fig. 50. — Trichonympha campanula Kof. and Swez. (After Kofoid and Swezy.) 



aggregate of kinetic elements of the cortical zone divides with it, 

 forming a mitotic figure with centrosomes, central spindle and astral 

 rays (Fig. 51). The connecting fibrils of the centrosomes, unlike 

 the centrodesmose in INIetazoa, remain outside of the nucleus (as 

 it does in many other flagellates) and is called the paradesmose by 

 Kofoid to distinguish it from the centrodesmose or central spindle. 

 Superficially, at least, this highly complicated type of mitotic figure 

 resembles the division figure of Noctihica miUaris in which the 

 kinetic elements are also extranuclear throughout the entire process. 

 Here, however, there is no such development of powerful motile 

 organoids and the division figure is relatively simple (Fig. 52). 

 As specially modified parabasal bodies, finally, may be included 



