364 Ciliophora 



would add a sixth family to Kahl's original five, but without eliminating 

 the need for further study of the remaining Frontoniidae. 



Family 1. CoJmilembidae. This family was erected for Cohnilembus 

 Kahl (102, 106; Fig. 7. 18, E). Lembadionella Kahl (106) was subsequently 

 referred to the family, and more recently, Anophrys Cohn (Fig. 7. 18, L) 

 has been added (155). The somatic cilia tion of these ciliates is complete 

 and rather vuiiform and the adoral ciliature consists of four membranes. 

 A paroral, or lateral, membrane extends along the right margin of the 

 elongated peristome. Three adoral membranes, which lie to the left in 

 the oral pouch of Tetrahymenidae (Fig. 7. 19, C), are shifted to the right 

 in Anophrys as a linear series parallel to the paroral membrane (Fig. 7. 

 18, L). There appears to be no row of somatic cilia ending at the poste- 

 rior margin of the oral cavity. In stomatogenesis during fission of 

 Anophrys (155), the adoral organelles of the posterior daughter are de- 

 rived from basal granules which undergo mvdtiplication at the base of the 

 paroral membrane. This type of stomatogenesis differs from that in the 

 Tetrahymenidae, as described below. 



Family 2. Frontoniidae. Although defining the family as one in which 

 the oral cavity does not open onto a clearly defined peristome, Kahl (102) 

 pointed out that the lack of information concerning adoral organelles 

 was responsible for much uncertainty in regard to the generic composi- 

 tion of the Frontoniidae. Later investigations have shown that Kahl's 

 uncertainty was justified. Removal of the Tetrahymenidae still leaves the 

 residual Frontoniidae a group probably in need of further subdivision. 



After elimination of certain ciliates more or less closely related to Tetrahymena 

 Furgason, the family includes the following genera: Aristerostoma Kahl (102), Bala- 

 nonema Kahl (102; Fig. 7. 18, C), Bizone Lepsi (102), Cardiostoma Kahl (102), Chas- 

 matostoma Engelmann (102), Cinetochilum Perty (102), Cryptochilidium Schouteden 

 (172; Fig. 7. 18, I), Cyrtolophosis Stokes (102), Dexiotrichides Kahl (102), Dichilum 

 Schewiakoff (102), Disematostoma Lauterborn (102; Fig. 7. 19, A), Epimecophrya Kahl 

 (106), Espejoia Biirger (66; Fig. 7. 19, I, J), Eurychilum Andre (102), Frontonia Ehrbg. 

 (102; Figs. 7. 18, A; 19, K, L), Frontoniella Wet/el (102). Homalogustra Kahl (102; Fig. 

 7. 18, B), Lanihornella Keilin (102), Leucophrydium Roux (102), Lembadion Perty 

 (102; Figs. 7. 18, J; 19, E), Malacophrys Kahl (102). Moiwrhihim Schewiakoff (102), 

 Platynematum Kahl (102; Fig. 7. 18, D), Pseudoglaucoma Kahl (102), Rhinodisculus 

 Mansfield (102), Saprophilus Stokes (102; Fig. 7. 18, F), Stegochilum Schewiakoff (102), 

 Stokesia Wenrich (219), Turaiiia Brodsky (102), Uvocentrum Nitzsch (77. 102; Figs. 7. 

 18, G; 19, F, G), Urojiema Dujardin (162; Fig. 7. 18, H), Uronemopsis Kahl (102), 

 Uropedalium Kahl (102), and Urozona Schewiakoff (102). 



Family 3. Tetrahymenidae. As shown by Furgason (69), the adoral 

 ciliature is composed of four membranes, three adoral ones lying to the 

 left in the oral pouch and a paroral membrane extending along the right 

 margin (Fig. 7. 19, C). Another feature is the presence of one or more 

 ciliary rows ("stomatogenous rows") which end at the posterior margin 

 of the oral pouch (Fig. 7. 20, H-L). In stomatogenesis the adoral mem- 



