T. F. GOREAU 281 



bonate. In corals, the coupling of the calcification reaction to 

 photosynthesis, though facultatixe, is almost certainly due to a 

 direct link \'ia a common metabolic pathway, rather than to 

 synthesis and diffusion of nutrients from the zooxanthellae to the 

 host. There can be no question that the great increase in rate and 

 efficiency of limestone secretion associated with photosynthesis 

 must, on a community level, be of decisive importance to the for- 

 mation, growth and maintenance of tropical coral reef ecosystems. 



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