174 LEECHES 



BDELLAMARIS Richardson 1953. Like Calliohdella or 

 Trachelohdella except annuli 12, diverticula absent. One sp. from 

 New Zealand. 



BRANCHELLION Savigny 1822 (Harding, 1910; Sukatschoff, 

 1912; Ingram, 1957). Suckers large, eyes 0-3 pairs, posterior 

 sucker containing many minute subsidiary suckers, constricted 

 clitellum partly covered by preputial fold of abdomen, abdomen 

 flattened with 31-33 pairs of lateral foliaceous branchiae (3 per 

 somite), annuli 3 (6), pulsatile vesicles 11 pairs, coelom typical, 

 diverticula fused, testes 5-6 pairs. At least 6 spp. mostly on 

 elasmobranchs. 



*Branchellion borealis Leigh- Sharpe (1933). 30 mm long, 

 31 pairs branchiae, no eyes, 4 tubercles dorsally on each of 4 

 anterior somites of trachelosome. Dorsally on Raja clavata. 

 English Channel. 



Branchellion ravenelii (Girard) (Meyer, 1941). 15 mm long, 

 31 pairs branchiae, 1 pair branching eye-spots. Ventrally on 

 Dasyatis hastatus, Amphotistius sabinus, Aetobatus freminvillii. 

 Florida and Carolina. 



Branchellion torpedinis Savigny. 50 mm long, 33 pairs branchiae. 

 On Rhinobatis thouin, Raja clavata^ Torpedo, Trygon pastinaca, 

 Labrus, Rhombus maximus. Mediterranean, North Sea, Ireland, 

 Senegal. 



CALLIOBDELLA Beneden and Hesse 1863 (Johansson, 1896; 

 Oka, 1910; Selensky, 1915; Leigh-Sharpe, 1914 and 1916). 

 Suckers discoid, eyes 0-2 pairs, posterior sucker broader than body, 

 abdomen more or less rounded, annuli 3 (6), pulsatile vesicles 11-13 

 pairs, coelom typical, diverticula more or less fused, testes 6 pairs, 

 rounded muscular organ connected with bursa. Several spp. 



*Calliobdella lophii Beneden and Hesse. 50 mm long, eyes 0, 

 posterior sucker 3^ X breadth of oral sucker, abdomen some- 

 what flattened and trachelosome well defined. On Lophius pisca- 

 torius. W. Europe. 



*Calliobdella nodulifera (Malm). 30 mm long, eyes 0, posterior 

 sucker 2 X breadth of oral sucker, abdomen rounded and 

 trachelosome ill defined. Externally on a variety of fish, especially 

 gadoids. W. Europe, Faroes, Iceland. 



Calliobdella punctata Beneden and Hesse. 20 mm long, eyes 



